Syspasis optica ( GRAVENHORST , 1829)

Riedel, Matthias, 2021, Revision of the Palaearctic species of Syspasis TOWNES, 1965 (Hymenoptera, Ichneumonidae, Ichneumoninae), Linzer biologische Beiträge 53 (2), pp. 775-810 : 790-791

Syspasis optica ( GRAVENHORST, 1829) (figs 9, 20, 42, 56)

Original: Ichneumon opticus GRAVENHORST, 1829: 131. L e c t o t y p e: (♁) [red] Lectotypus Ichneumon opticus GRAVENHORST 1829 ♁ (Wroclaw), type studied. syn. nov. Ichneumon simulosus THOMSON, 1886: 16. L e c t o t y p e: (♀)Hbg; simulosus (Lund), lectotype herewith designated, type studied. syn. nov. Ichneumon rimulosus DALLA TORRE, 1902. M a t e r i a l e x a m i n e d: 3♀♀, 9♁♁; Belgium *, Finland, Germany, Kazakhstan, Poland,

Russia (Dnjepr, Caucasus).

D e s c r i p t i o n: ♀: Body length 9-10 mm. Flagellum with 31-32 flagellomeres, slightly lanceolate; 1 st flagellomere 3.0× and 2 nd flagellomere 2.5-2.7× longer than wide, combined length 0.70-0.74× eye length; 9 th flagellomere square; widest flagellomeres 1.3× wider than long. Temple strongly and almost linearly narrowed behind eye. OED 1.5× ocellar diameter. Frons transversely rugose-punctate or with superficial punctures. Face and clypeus punctate and granulate. Gena with superficial punctures, 1.0× as wide as eye. MI 0.9. Mesosoma covered with dense whitish hairs. Side of pronotum punctate dorsally and striate in ventral 2/3. Notaulus impressed frontally. Mesoscutum with superficial punctures, granulate. Mesopleuron densely punctate, with striation caudally and dorsally. Metapleuron punctate, sometimes rugose. Scutellum slightly wider than long, punctate, with lateral carina in basal 0.1-0.3. Area basalis bulged forward medially, without distinct median tubercle. Area superomedia large, heart-shaped, about as long as wide, almost smooth; anterior transverse carina caudal to its middle. Area externa with superficial punctures, granulate. Area petiolaris rugose-punctate, without lateral carina. Hind coxa densely pilose, with large, diffuse apico-ventral scopa. Hind femur with scattered punctures ventrally, 3.7-4.3× longer than wide. Areolet pentagonal, frontal distance between veins 2rs-m and 3rs-m 2-3× their width; vein 2m-cu slightly distad its middle. Vein 1cu-a interstitial or slightly postfurcal. Metasoma oxypygous. Lateral field of postpetiolus 0.5-1.0× as wide as median field, the latter with c. 8 coarse longitudinal striae. Thyridium transverse, 1.8-2.5× as wide as the interval between thyridia. 2 nd tergite 0.75- 0.80× as long as wide. 2 nd and 3 rd tergites with very superficial punctures, granulate; 2 nd and sometimes 3 rd tergites ± strongly and widely striate. Ovipositor sheath moderately extending behind metasomal apex (by 0.5-0.7× length of 7 th tergite).

Color: black. Stripes or rings on flagellomeres 7/8-10/11 yellowish. 1 st to 3 rd tergites reddish or brownish, 4 th tergite sometimes reddish basally, following tergites black. Coxae and trochanters black; femora black or reddish, if reddish, then ± infuscate apically; tibiae red or darkened, fore and mid tibiae ochreous frontally; tarsi dark. Pterostigma pale brownish. ♁: Body length 11-12 mm. Flagellum with 32-33 flagellomeres; 1st flagellomere 2.5×

and 2 nd flagellomere 2.1-2.2× longer than wide, combined length 0.71-0.72× eye length. Tyloids on flagellomeres 7/9-15/18, linear, maximally 0.7-0.8× as long as the flagellomeres. Temple moderately or strongly and roundly narrowed behind eye. OED 1.5× ocellar diameter. Gena slightly concave ventrally, 0.85-0.9× as wide as eye. MI 0.6. Mesosoma with pale brownish hairs. Scutellum about as long as wide, punctate and shortly striate apically, with lateral carina in basal 0.3-0.7. Area superomedia wider than long, hexagonal or horseshoe-shaped, fine rugulose. Area externa with scattered punctures, almost smooth. Hind femur densely punctate, 3.9-4.3× longer than wide. Fore metatarsus without apical tooth. Wings as in ♀. Lateral field of postpetiolus rugose, 0.8- 0.9× as wide as median field, the latter with c. 7-8 coarse longitudinal striae. Gastrocoelus distinct, with longitudinal ridges. Thyridium oblique, 2.0-2.8× wider than the interval between thyridia. 2 nd tergite 0.85-1.05× as long as wide. 2 nd to 4 th tergites with fine superficial rugose-punctation, dull, aciculate medially. Aciculation on 2 nd (and sometimes 3 rd tergite) tergites wide (covering about 2/3 of tergal width).

Color: black. Facial orbit, sometimes frontal orbit in ventral half, outer orbit, spot on vertex, sometimes scape ventrally, collar (sometimes weak), hind edge of pronotum, often spot on subtegular ridge yellow. Tergites black, usually narrowly reddish at apical margins, but sometimes 1 st to 4 th tergites dark reddish. Coxae and trochanters blackish. Legs otherwise brownish or reddish; fore and mid legs paler frontally; hind tibia apically and hind tarsus completely blackish. Pterostigma reddish-brown.

Hosts: unknown.