Syspasis albiguttata ( GRAVENHORST , 1820)

Riedel, Matthias, 2021, Revision of the Palaearctic species of Syspasis TOWNES, 1965 (Hymenoptera, Ichneumonidae, Ichneumoninae), Linzer biologische Beiträge 53 (2), pp. 775-810 : 779-780

Syspasis albiguttata ( GRAVENHORST, 1820) (figs 1, 16, 24, 34, 48)

Original: Ichneumon albiguttatus GRAVENHORST, 1820: 280.

H o l o t y p e: (♀) Oe. (Wroclaw), type not studied.

syn. Ichneumon multicinctus GRAVENHORST, 1829: 453, synonymized by WESMAEL (1857).

syn. Amblyteles albostriata TISCHBEIN, 1881: 185 , synonymized by CONSTANTINEANU (1959).

syn. Ichneumon ebeninus BERTHOUMIEU, 1895: 284, synonymized by DILLER & HORSTMANN (1997). syn. nov. Ctenichneumon maruyamensis UCHIDA, 1930: 97 , high resolution fotos studied.

M a t e r i a l e x a m i n e d: 80♀♀ and 11♁♁ from Austria, Belgium, Czech Republic, France, Germany, Japan *, Poland, Russia (Far East), Spain, Switzerland, Turkey *.

D e s c r i p t i o n: ♀: Body length 9-14 mm. Flagellum slightly lanceolate, with 35-39 flagellomeres; 1 st flagellomere 2.7-2.8× and 2 nd flagellomere 2.1-2.4× longer than wide; widest flagellomeres 1.2× wider than long. Temple moderately and roundly narrowed behind eye. OED 1.0× ocellar diameter. Frons granulate and coarsely but superficially punctate dorsally, ± smooth ventrally. Face and clypeus granulate and punctate. Gena 0.65× as wide as eye. MI 1.0. Mesosoma covered with short whitish hairs. Side of pronotum finely punctate dorsally and striate ventrally. Mesoscutum finely granulate and densely but superficially punctate, ± dull. Notaulus indistinctly impressed on mesoscutum. Mesopleuron striate-punctate. Metapleuron punctate, with ± rugose margins; juxtacoxal carina present. Scutellum slightly elevated, wider than long, punctate, with lateral carina in basal 0.4-0.5. Area basalis with small median tubercle. Area superomedia horseshoe- or sickle-shaped, slightly wider than long, finely rugose or aciculate; anterior transverse carina present, reaching area superomedia slightly posterior to its middle. Area externa smooth, superficially punctate. Area petiolaris rugose-punctate, lateral carina indistinct. Hind coxa densely punctate, without scopa. Hind femur densely punctate, 3.9-4.0× longer than wide. Areolet pentagonal, frontal distance between veins 2rs-m and 3rs-m 1-2× their width. Vein 2m-cu reaching areolet in its middle. Vein 1cu-a slightly postfurcal. Metasoma strongly oxypygous. Postpetiolus strongly widened. Lateral field finely rugose, 0.6× as wide as median field; median field with c. 11 fine striae and partly punctate. Thyridium oblique, 0.8-1.0× as wide as the interval between thyridia. 2 nd tergite 0.80-0.85× as long as wide, densely punctate and granulate, dull, often ± aciculate medially. 3 rd tergite with superficial punctures; 4 th tergite almost smooth, finely granulate. Ovipositor sheath slightly extending behind metasomal apex (by 0.4-0.8× length of 7 th tergite).

Color: black. Ivory coloration variable: usually 8 th to 12/13 th flagellomeres with ivory rings or stripes; frontal orbit (widened dorsally), outer orbit dorsally, often collar and hind edge of pronotum, often spot on subtegular ridge and apical stripe on scutellum, often median spot on hind margin of postpetiolus, often median spots or stripes on hind margins of 3 rd /4 th to 5 th tergites, fore femur and tibia frontally, mid and hind femora apically ivory. Legs otherwise brown or black. Pterostigma brownish, paler centrally.

♁: Body length 10-14 (18) mm. Flagellum with 36-39 flagellomeres; 1 st flagellomere 2.5-2.8× and 2 nd flagellomere 2.0× longer than wide. Tyloids on flagellomeres 9/11-19, linear, maximally 0.9× as long as their flagellomeres. Temple moderately and roundly narrowed behind eye. OED 0.8-1.0× ocellar diameter. Occiput rather deeply excavated medially. Frons with superficial punctures dorsally, ± smooth ventrally. Face and clypeus punctate and granulate. Gena 0.8× as wide as eye. MI 0.3-0.4. Pronotum with fine superficial punctures dorsally, with largely smooth surface, with fine striae ventrally. Scutellum wider than long, with lateral carina in its basal 0.6-0.8. Hind femur 3.8-4.2× longer than wide. Lateral field of postpetiolus 0.7× as wide as median field, the latter aciculate and punctate. Thyridium about as wide as the interval between thyridia. 2 nd and 3 rd tergites punctate, partly granulate, ± aciculate medially.

Color: black. Palps, mandible, clypeus, face, wide frontal orbit, outer orbit, scape ventrally, collar and hind edge or upper margin of pronotum, usually ventral edge of pronotum, lateral and apical margins of scutellum, subtegular ridge, often spot on tegula, apical margins of 1 st to 2 nd /3 rd tergites, apical spots on 3 rd /4 th to 5 th /6 th tergites, spots on fore and mid coxae and trochanters, often spot on hind coxa, fore and mid legs frontally, hind femur in apical 0.2, often apical spot on hind tibia, sometimes spot on mesopleuron, hind edge of area dentipara and postscutellum yellow.

Hosts: Sphrageidus similis (FUESSLY, 1775) (Lep., Noctuidae ), Calliteara pudibunda (LINNAEUS, 1758) (Lep., Erebidae ) ( RASNITSYN & SIYTAN 1981).

T a x o n o m i c a l r e m a r k: Thetypeof Ctenichneumon maruyamensis UCHIDA is larger (body length 18 mm) than other ♁♁ of S. albiguttata , but structure and coloration are typical. I saw another Japanese ♁ of S. albiguttata (from Nagano Prefecture / Japan in Kanagawa Museum) which confirms the occurrence of this species in Japan, but is smaller with a body length of 11 mm. I synonymize both species here.