Syspasis TOWNES, 1965

Riedel, Matthias, 2021, Revision of the Palaearctic species of Syspasis TOWNES, 1965 (Hymenoptera, Ichneumonidae, Ichneumoninae), Linzer biologische Beiträge 53 (2), pp. 775-810 : 777-778

Key for the Palaearctic species of Syspasis TOWNES

(modified after RASNITSYN & SIYTAN 1981)

Females:

1 Mesosoma entirely black (figs 34, 36, 39). Metasoma ± red...............................................2

- Mesosoma with ivory stripes (figs 35, 37, 38) ....................................................................4

2 Area basalis without distinct median tubercle (fig. 36). Scutellum not carinate. Apical tergites black. Hind femur mainly black. Hind tibia with 2 external denticular spurs. Flagellum with 28-30 flagellomeres. Palaearctis................ S. haesitator (WESMAEL, 1845)

- Area basalis with strong median tubercle (fig. 41). Scutellum ± carinate laterally (fig. 41). Hind tibia with multiple external denticular spurs. Hind femur red. Flagellum with at least 32 flagellomeres. .....................................................................................................3

3 Mid and hind tibiae mainly black. Flagellum with 37 flagellomeres. Hind femur slender, 4.4× longer than wide. Malar space 1.1× as long as width of mandibular base (fig. 8). Areolet quadrilateral: veins 2rs-m and 3rs-m meeting frontally. Kyrgyzstan........... ......................................................................................................... S. nigrotibialis nov.sp.

- Mid and hind tibiae mainly red. Flagellum with 32-33 flagellomeres. Hind femur stout, 3.8-4.2× longer than wide. Malar space 0.9-1.0× as long as width of mandibular base. Areolet ± pentagonal: frontal distance between veins 2rs-m and 3rs-m 2× their width. Middle Asia....................................................................................... S. nigrithorax nov.sp.

4 Metasoma entirely red OR at least 2 nd to 4 th tergites red. Prescutellar carina usually ivory. Outer orbit with long ivory stripe. Pterostigma dark. Mesoscutum sometimes with two paramedian ivory stripes. Body length 10-12 mm ................................................5

- Metasoma usually mainly black, at most basal tergites reddish. Prescutellar carina black. Outer orbit often without long ivory stripes. ............................................................7

5 Apical margin of clypeus with small median tooth (fig. 15). Hypostomal carina lamelliform elevated; genal carina ± obsolete ventrally (fig. 15). Hind coxa without scopa. West Palaearctis. ........................................... S. leucolomia ( GRAVENHORST, 1829)

- Apical margin of clypeus without median tooth. Hypostomal carina narrow; genal carina complete ventrally, reaching hypostomal carina.......................................................6

6 Apical tergites red. Temple strongly narrowed behind eye (fig. 25). Palaearctis. ................. ........................................................................................... S. carinator ( FABRICIUS, 1798)

- 5 th to 7 th tergites black. Temples shortly parallel behind eye, moderately narrowed apically (fig. 29). Siberia................................................................ S. nigroapicalis nov.sp.

7 Thyridium about as wide as the interval between thyridia (fig. 48). Usually hind edge of scutellum and flagellar ring, sometimes postpetiolus and 3 rd to 5 th tergites apically with ivory spots. Temple strongly narrowed behind eye (fig. 24). Pterostigma brown, paler centrally. Mesoscutum densely punctate and granulate, dull. 1 st flagellomere 2.7- 2.8× longer than wide. Body length 10-15 mm. Palaearctis. ................................................. .................................................................................. S. albiguttata ( GRAVENHORST, 1820)

- Thyridium much wider than the interval between thyridia (fig. 49). Metasoma without ivory spots. ..........................................................................................................................8

8 Mesosoma including scutellum completely black (figs 42, 43). Flagellum with at most 36 flagellomeres. .................................................................................................................9

- Side of scutellum ± ivory (figs 38, 44). Flagellum with at least 34 flagellomeres. ...........11

9 2 nd and 3 rd tergites superficially punctate, at most narrowly (less than 0.5× tergal width) and finely striate medially, black (fig. 57). Hind coxa without scopa. Body length 10 mm. Palaearctis.. ........................................................ S. puerulus ( KRIECHBAUMER, 1890)

- 2 nd and 3 rd tergites widely (more than 0.5× of tergal width) with distinct longitudinal striae or rugae, usually ± red (figs 56, 60). Hind coxa with or without scopa. ..................10

10 Hind coxa with scopa. 1 st flagellomere 3.0× longer than wide. Temple distinctly and linearly narrowed behind eye. Frontal orbit without ivory stripe. Metasoma entirely black OR 1 st to 3 rd tergites ± reddish. Body length 9-11 mm. West Palaearctis and Kazakhstan. ...................................................................... S. optica ( GRAVENHORST, 1829)

- Hind coxa without distinct scopa. 1 st flagellomere 2.0× longer than wide. Temple slightly and roundly narrowed behind eye. Frontal orbit with ivory stripe. Postpetiolus and 2 nd to 3 rd tergites red. Body length 11 mm. North China. ................. S. sinensis nov.sp.

11 Pterostigma, femora and tibiae red. Temple wide, slightly narrowed apically. Outer orbit black. Flagellum with 35-38 flagellomeres, 1 st flagellomere long, 3.3-3.8× longer than wide. Basal tergites sometimes ± reddish. Body length 10-14 mm. Palaearctis ............ .................................................................................. S. scutellator ( GRAVENHORST, 1829)

- Pterostigma and often legs ± brown or black. Temple narrower, usually ± narrowed behind eye.........................................................................................................................12

12 1 st flagellomere 2.5-2.8× longer than wide. Hind coxa with distinct scopa. Hind femur stout, 3.4-3.8× longer than wide. Inner orbit with small ivory spot opposite antennal groove. Hind edge or upper margin of pronotum yellowish. Body length 9-13 mm. Holarctis. .................................................................................. S. tauma ( HEINRICH, 1951)

- 1 st flagellomere 2.9-3.5× longer than wide. Hind coxa with weak and indistinct scopa. Hind femur slenderer, 3.9-4.3× longer than wide. Frontal orbit with ivory stripe. Pronotum without ivory spots. Body length 12-15 mm. Palaearctis...................................... .............................................................................................. S. lineator (FABRICIUS, 1781)