Coptoprepes nahuelbuta, RAMÍREZ, 2003

RAMÍREZ, MARTÍN J., 2003, The Spider Subfamily Amaurobioidinae (Araneae, Anyphaenidae): A Phylogenetic Revision At The Generic Level, Bulletin of the American Museum of Natural History 2003 (277), pp. 1-262 : 85-87

Coptoprepes nahuelbuta , new species Figure 38

TYPES: Female holotype and male paratype from Chile, Región IX, Malleco province, Nahuelbuta Natl. Park, FITS, 1200 –1500 m, Nothofagus / Araucaria forest, ca. 38°S, 73°W, 9.XII.1984 – 17.II.1985, S. and J. Peck, deposited in AMNH.

ETYMOLOGY: The specific name is a noun in apposition, referring to the area where this species lives.

DIAGNOSIS: Distinguished from other Coptoprepes by having a projecting secondary conductor, articulated and heavily sclerotized, and an epigynal median field hidden in the epigastric furrow.

FEMALE (holotype): Total length 5.30. Carapace length 2.03, width 1.43, wider on leg II. Length of tibia/metatarsus: I, 0.92/0.84; II, 0.83/0.80; III, 0.72/0.89; IV, 1.10/1.07. Palpal tarsus length 0.57. Chelicerae with five teeth on promargin, seven denticles on retromargin. Sternum length 1.08, width 0.87. Metatarsi III and IV with preening comb. Spines: leg I, femur d 1–1–1, p 2ap; tibia v 2–2–0; metatarsus v 2bas. II, femur d 1–1– 1, p d1ap; tibia v r1­ r1–2; metatarsus = I. III, femur d 1–1–1, p 0­ d1­ d1 or d1ap, r d1ap or 0; patella r 1; tibia v p1­ p1–2, p 1­ d1­ 1­ 0, r d1–1, d r1bas; metatarsus v 2–0­ comb, p and r d1–1–1, d 0­ p1–2. IV, femur d 1–1–1, p and r d1ap; patella r 1; tibia v p1–2–2, p and r 1­ d1­ 1­ 0, d r1bas; metatarsus v 2–2­ comb, p and r d1–1–1, d 0–2–2. Dorsal bristles as in C. flavopilosus . Abdomen (with cuticle partially detached) length 3.55. Color: holotype quite faded; other specimens with carapace, legs, sternum, mouthparts brown, abdomen dark grayish, dorsum paler, cardiac area dark. Epigyne (fig. 38F– H) displaced posteriorly, in ventral view only visible as elevation of epigastrium. Lateral lobes separate in posterior view. Copulatory ducts contorted, asymmetrical, spermathecae contiguous, lumen small.

MALE (paratype): Total length 4.12. Carapace length 1.90, width 1.37. Length of tibia/ metatarsus: I, 1.23/1.09; II, 1.04/0.99; III, 0.81/1.00; IV, 1.17/1.47. Chelicerae smaller than those of female. Sternum length 1.06, width 0.78. Spines as in female, except: leg I, tibia v 2–2–2. II, tibia p 0–1. Abdomen (badly preserved) length ca. 2.00. Color as in female. Palp (fig. 38A–D): femur short; tibia very short, width/length 1.39, RTA long, flattened, distally bent dorsally. Cymbium relatively large, with deep apical retrolateral notch where median apophysis and secondary conductor fit; cymbial conductor wide. Tegulum basal. Sperm duct with pronounced loop at dorsal anterior margin, between secondary conductor and median apophysis. Embolus with basal process ample, flattened, rounded. Median apophysis apical, sinuous, forming right angle, apex very thin. Primary conductor with deep canal, apical portion projecting as straight, heavily sclerotized prong, where canal ends. Secondary conductor conspicuous, heavily sclerotized, almost totally surrounded by membranous area (fig. 38B). Paramedian apophysis absent, or represented only by a longitudinal ridge close to base of median apophysis; rounded, ventral protuberance, may also be part of paramedian apophysis.

VARIABILITY: Females spines: I, tibia v 2– 2­ p1 or 2–2–2. II, tibia v r1­ r1­ p1. IV, metatarsus d 0­ p1–2.

NATURAL HISTORY: Unknown. All specimens were collected in pitfall traps or in leaf litter.

DISTRIBUTION: Forests in southern Chile, in Malleco and Aisén provinces, probably also in intermediate localities.

OTHER MATERIAL EXAMINED: CHILE: Región IX (Araucanía): Malleco: Same data as types, 23 3♀ (AMNH); 17 km W Angol, 800 m, FIT, mixed Nothofagus , 8.XII.1984 – 16.II.1985, S. and J. Peck, 13 (AMNH); Monumento Natural Contulmo, 11.XII.1984 – 13.II.1985, S. and J. Peck, 13 1♀ (AMNH); Nahuelbuta Natl. Park, 1250 m, 19.XI.1981, N. Platnick and T. Schuh, mossy forest litter, Nothofagus , Araucaria , 1♀ (AMNH). Región XI (Ibáñez del Campo): Aisén: 102 km S Puerto Puyuguapi, 220–270 m, burned forest, 19.I.1986, N. Platnick, P. Goloboff, T. Schuh , 2♀ (AMNH).

Figures

Fig. 38. Coptoprepes nahuelbuta, n. sp. A. Male palp, ventral view (paratype). B. Same, copulatory bulb, apical view. C. Same, ventral­apical view. D. Same, retrolateral view. E. Same, palp, retrolateral view. F. Epigyne, ventral view (holotype). G. Same, posterior view. H. Same, dorsal view, cleared. Scale bar = E, 0.4 mm; H, 0.15 mm, all other, 0.2 mm. (C1 = primary conductor; C2 = secondary conductor; E = embolus; MA = median apophysis; PBE = process on base of embolus.)