Aysenia segestrioides, RAMÍREZ, 2003

RAMÍREZ, MARTÍN J., 2003, The Spider Subfamily Amaurobioidinae (Araneae, Anyphaenidae): A Phylogenetic Revision At The Generic Level, Bulletin of the American Museum of Natural History 2003 (277), pp. 1-262 : 62-64

Aysenia segestrioides , new species Figure 23

TYPE: Male holotype from Chile, Región X, Valdivia province, Las Lajas, W La Unión, ca. 40°46̍S, 73°42̍W, 19 – 20.XI.1990, L. Peña, deposited in AMNH.

ETYMOLOGY: The specific name refers to the striking similarity with segestriid spiders, because of the elongate body and the third leg directed forward.

DIAGNOSIS: Easily distinguished by having a huge primary conductor occupying most of the male copulatory bulb, and greatly coiled female copulatory ducts.

FEMALE (Valdivia, AMNH): Total length 8.10. Carapace elevated, cephalic area flat; length 2.93, width 1.57, wider behind chelicerae. Length of tibia/metatarsus: I, 0.90/0.79; II, 0.67/0.77; III, 0.62/0.66; IV, 1.02/0.99. Palpal tarsus length 0.69. Chelicerae very strong, with two teeth on retromargin, fang short, thick (fig. 23G). Sternum length 1.27, width 0.73. Spines: femora I–IV d 1–1–1. Leg I, tibia v r1­ r1–2, p v1–1–0 (these are the ventrals displaced); metatarsus v 2–0. II, tibia v r1­ r1­ r1, p 0­ d1; metatarsus v 2bas, p 1. III, tibia v r1ap, p and r d1; metatarsus v 2–0–2, p 0­ d1–1, r 1ap, d 2ap. IV, tibia v p1­ p1–2; metatarsus v 2­ p1–2, p and r 1ap, d r1ap. Scopulae on tarsi I and II, and metatarsus II, denser on prolateral faces. Leg III oriented forward (fig. 23F). Femora I–III narrow, IV very strong. Abdomen length 5.30, width 2.33, spiracle–epigastrium 3.13, spiracle–spinnerets 0.30. Color: carapace dark brown, ocular area almost black. Legs brown, I and II darker. Sternum uniform brown, endites and labium dark brown. Abdomen grayish brown with cream pattern, venter grayish uniform. Epigyne (fig. 23D, E): lateral lobes well sclerotized, separate, elevated above median field. Copulatory ducts very long, coiled.

MALE (holotype): Total length 5.00. Carapace length 2.40, width 1.37 (fig. 23A). Length of tibia/metatarsus: I, 1.05/1.00; II, 1.05/0.97; III, 0.73/0.82; IV, 1.05/1.00. Chelicerae smaller than those of female. Sternum length 1.08, width 0.68. Spines as in female, except: leg I, tibia v r1­ r1–2 or r1­ r1­ p1, p d1­ 0­ d1­ 0; metatarsus v 2–2–0, p 0–1­ d1, d p1. II, tibia v r1­ r1–2; metatarsus v 2bas, p 0–1­ d1. III, tibia p d1 or d1­ d1–0. IV, metatarsus p 0–1–1. Abdomen length 2.60, width 1.30, spiracle–epigastrium 1.20, spiracle– spinnerets 0.23. Color as in female. Palp (fig. 23B, C): tibia short, width/length 0.93, RTA sharp, long. Cymbial conductor wide. Copulatory bulb extremely modified, distal region occupying ventral face, primary conductor covering tegulum. Sperm duct with two conspicuous loops at anterior dorsal margin, one at base of secondary conductor. Embolus very long, thin, basal process rounded. Median apophysis apical, hooked, tip pointing basally and retrolaterally. Primary conductor huge, crescent­shaped, projecting at both ends; canal area covered by thin projections. Secondary conductor flattened, partly fused to anterior margin of tegulum, without canal. Paramedian apophysis with three cusps, two heavily sclerotized, close to median apophysis, third one less sclerotized, flattened, close to primary conductor.

NATURAL HISTORY: Unknown. The prolateral displacement of both ventral spines on first tibiae, and of scopulae of first and second legs, suggest further adaptations for living in tubes.

DISTRIBUTION: Known only from Valdivia province.

OTHER MATERIAL EXAMINED: CHILE: Región X (Los Lagos): Valdivia: Valdivia , no date, collection E. Simon, 13 ( MHNP 18235 ) ; Valdivia, 12.X.1976, E. Krahmer, 1♀ ( AMNH) .

Figures

Fig. 23. Aysenia segestrioides, n. sp., male holotype, female from Valdivia (AMNH). A. Male. B. Male palp, ventral view. C. Same, detail retrolateral­ventral. D. Epigyne, ventral view. E. Cleared epigyne, dorsal view. F. Female. G. Female chelicera, ventral view. Scale bars = A, F, 1 mm; B–E, 0.2 mm; G, 0.5 mm. (FD = fertilization duct.)