Euphthiracarus ornatus, : Niedbala & Liu, 2018

Niedbała, Wojciech & Liu, Dong, 2023, Systematic, synonymic and biogeographical list of ptyctimous mites (Acari, Oribatida) in the world (1799 – 2022), Zootaxa 5265 (1), pp. 1-442 : 159-163

Neotropical, Mesoamerica ornatus Phthiracarus Mahunka, 1991

Phthiracarus (Archiphthiracarus) ornatus Mahunka, 1991

Phthiracarus ornatus: Niedbała 2000, 2004c , Niedbała & Liu 2018.

Phthiracarus (Archiphthiracarus) : Subías 2022.

Orient: Malaysia (Sabah) ( Balogh & Balogh 2002, Mahunka 1991b, Niedbała 2000, 2002, 2004c, Subías 2022).

Oriental, Malaysia, endemic ortizi Steganacarus (Rhacaplacarus) Pérez-Íñigo, 1969

Steganacarus ortizi: Pérez-Íñigo, 1969a

Steganacarus ortizi: Pérez-Íñigo 1969c , Balogh & Mahunka 1980 a, 1983.

Steganacarus (Rhacaplacarus) ortizi: Niedbała 1986b, 1992 , 2011b, 2012b, Bernini & Magari 1993, Avanzati et al. 1995, Niedbała & Liu 2018.

Steganacarus (Steganacarus) ortizi: Bernini & Avanzati 1986 .

Rhacaplacarus (Rhacaplacarus) ortizi: Subías 2004 .

Hoplophorella (Rhacaplacarus) ortizi: Subías 2022 .

Palaearctic: England ( Jersey Island), Spain, southern France, Italy, Madeira Islands ( Avanzati et al. 1995, Balogh & Mahunka 1980 a, 1983, Bernini & Avanzati 1986, Bernini & Magari 1993, Niedbała 1986b, 1992, 2011b, 2012b, Pérez-Íñigo 1969a, c, Pigliucci et al. 1990, Subías 2022).

Western Palaearctic, reported from Western Europe and Atlanto-Mediterranean region otagoensis Notophthiracarus Niedbała, 2016

Notophthiracarus otagoensis: Niedbała & Ermilov 2016 , Niedbała & Liu 2018.

Hoplophthiracarus (Notophthiracarus) otagoensis: Subías 2022 .

Australasia: New Zealand ( Niedbała & Ermilov 2016, Subías 2022).

Australasian, New Zealand, endemic otaheitensis Acrotritia ( Hammer, 1972)

Rhysotritia ardua var. otaheitensis Hammer, 1972

Rhysotritia otaheitensis: Niedbała 1998b .

Acrotritia otaheitensis: Niedbała & Penttinen 2007 , Liu & Zhang 2014a, Liu 2015 d, Niedbała & Liu 2018.

Australasia: Tahiti, Tonga ( Balogh & Balogh 2002, Hammer 1972, 1973, Niedbała 1998b, 2012a, Niedbała & Penttinen 2007, Subías 2022).

Australasian, Pacific Islands

Note. Subías (2022) considered this species as congeneric with A. curticephala ( Jacot, 1938) . See note of A. curticephala .

othneios Atropacarus (Hoplophorella) Niedbała et Starý, 2014 synonym of Atropacarus (Hoplophorella) mahunkai Niedbała et Starý, 2013

othneios Plonaphacarus Niedbała, 2022

Hoplophthiracarus (Hoplophthiracarus) othneios: Subías 2022 .

Palaearctic: Nepal ( Niedbała & Ermilov 2022a).

Palaearctic, Nepal, endemic

Note. This species is very similar to P. martensi Niedbała, 1914 but differs in the lack of lateral carinae of prodorsum, formula of genital setae is 6(4+2): 3 (versus 7 (4+3): 2) and setae d of femora I spiniform (versus distal end of femora I with distinct spines).

otwayensis Notophthiracarus Niedbała, 2017

Notophthiracarus otwayensis: Niedbała & Szywilewska-Szczykutowicz 2017 , Niedbała & Liu 2018, Niedbała et al. 2021.

Hoplophorella (Hoplophorella) otwayensis: Subías 2022 .

Australasia: Australia (Victoria) (Niedbała et al. 2021, Niedbała & Szywilewska-Szczykutowicz 2017, Subías 2022).

Australasian, Australia, endemic pachetos Arphthicarus Niedbała, 2008

Arphthicarus pachetos: Niedbała & Liu 2018 .

Hoplophorella (Hoplophorella) pachetos: Subías 2022 .

Orient: Malaysia (Sabah) (Liu et al. 2013, Niedbała 2008b, Subías 2022).

Oriental, Malaysia, endemic pachys Phthiracarus Niedbała, 2015

Phthiracarus pachys: Niedbała & Liu 2018 .

Phthiracarus (Archiphthiracarus) pachys: Subías 2022 .

Palaearctic: Spain (Niedbała 2015b, Subías 2022).

Palaearctic, Spain, endemic paeneminima Rhysotritia Walker, 1965 synonym of Microtritia simplex ( Jacot, 1930)

pakistanensis Euphthiracarus (Euphthiracarus) Hammer, 1977

Euphthiracarus pakistanensis Hammer, 1977

Euphthiracarus (Euphthiracarus) pakistanensis: Niedbała 2000 , Subías 2004, Liu 2021b.

Euphthiracarus pakistanensis: Niedbała 2004c, 2011b , 2012b, Niedbała & Liu 2018.

Euphthiracarus meghalayensis Sanyal, 1988 syn. by Niedbała 2004c.

Euphthiracarus (Euphthiracarus) meghalayensis: Subías 2022 .

Palaearctic: Pakistan, India (Arunachal Pradesh, northern part), Russia (Far East), China (Gansu) ( Hammer 1977, Liu 2021b, Niedbała 2000, 2004c, 2011b, 2012b, 2015b, Niedbała & Ermilov 2013b, Subías 2022). Eastern Palaearctic, in Prooriental and Amurian areas pakistanensis Hoplophthiracarus ( Hammer, 1977)

Hoplophorella pakistanensis Hammer, 1977

Hoplophthiracarus pakistanensis: Niedbała 1984b, 1986b , 1992, 2000, 2011b, 2012b, Subías 2004, Niedbała & Liu 2018.

Hoplophorella (Rhacaplacarus) pakistanensis: Subías 2022 .

Palaearctic: Pakistan, India (northern part), Vietnam ( Balogh & Balogh 2002, Hammer 1977, Niedbała 1984b, 1986b, 1992, 2000, 2011b, 2012b, Subías 2022).

Southern Palaearctic, distributed in Kashmir-Pakistanian region, Vietnam palaciosi Prototritia Mahunka et Mejia-Recamier, 1998

Prototritia palaciosi: Niedbała 2004a , b, 2008e, Niedbała & Liu 2018.

Prototritia (Prototritia) palaciosi: Subías 2022 .

Neotropic: Mexico, Panama (Mahunka & Mejia-Recamier 1988, Niedbała 2004a, b, 2008e, Subías 2022). Neotropical, Mesoamerica pallens Atropacarus (Hoplophorella) (Niedbała, 1983)

Hoplophorella pallens Niedbała, 1983

Atropacarus (Hoplophorella) pallens: Niedbała 1986b, 1992 , 2001 a, Niedbała & Liu 2018. Hoplophorella (Hoplophorella) pallens: Subías 2022 .

Afrotropic: Mali, Togo, Kenya ( Balogh & Balogh 2002, Niedbała 1983i, 1986b, 1992, 2001a, Subías 2022).

Afrotropical, broad, discontinuous

Notes. This species does not occur in the Mesoamerica as erroneously reported in Niedbała (1992). Subías (2022) considered this species as congeneric with Atropacarus (Hoplophorella) hamatus (Ewing, 1909) . However, both species have different shape of notogastral setae.

pallidus Phthiracarus Feider et Suciu, 1958

Phthiracarus pallidus: Krivolutsky 1975 , Balogh & Mahunka 1980 a, 1983, Niedbała 1986b, 1992, 2011b, 2012b, Sergienko 1989b, Niedbała & Liu 2018.

Phthiracarus occultus Niedbała, 1981 syn. by Niedbała 2011b.

Phthiracarus occultus: Niedbała 1986b, 1992 , 2008f, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008.

Phthiracarus (Archiphthiracarus) occultus: Subías 2004 .

Palaearctic: Poland, Romania, Ukraine ( Balogh & Mahunka 1980a, Feider & Suciu 1958b, Mahunka et al. 2013, Niedbała 1981a, 1986b, 1992, 2008f, 2011b, 2012b, 2015b, Olszanowski et al. 1996, Subías 2022). Western Palaearctic, known from Central and Eastern Europe

Note. Subías (2004) recognized this species as congeneric with Phthiracarus compressus Jacot 1930 .

However, P. pallidus differs from P. compressus by the length of notogastral setae c 1 <1/2 c 1 - c 1 (versus c 1>1/2 c 1 - c 1 in P. compressus ), implantation of setae c 1 almost at anterior margin (versus remote from margin) and presence of lyrifissures ips (versus absent).

palpalis Protoplophora Berlese, 1910 s

Prothoplophora palpalis (lapsus linguae): Lombardini 1936.

Protoplophora palpalis: Hammen 1959 , Krivolutsky 1975, Subías & Pérez-Íñigo 1978, Balogh & Mahunka 1980 a, 1983, Mahunka & Mahunka-Papp 1995, Niedbała 2002a, 2004b, 2011b, 2012b, Niedbała & Liu 2018, Subías 2022.

Protoplophora sp. : Walter & Norton 1985, Marshall et al. 1987.

Nearctic: USA (California). Palaearctic: Spain, Italy, Sicily ( Balogh & Mahunka 1980 a, 1983, Berlese 1910, Bernini 1983, Bernini & Arcidiacono 1985, Castagnoli & Pegazzano 1985, Hammen 1959, Mahunka & Mahunka-Papp 1995, Marshall et al. 1987, Minguez-Martinez 1981, Niedbała 2002a, 2004b, Subías 1980, 2022). Doubtful findings: Middle Asia (Krivolutsky 1966), Tunisia ( Mahunka 1980), South Australia ( Lee 1981).

Holarctic, only in mediterranean parts (Californian and Mediterranean subregions)

paludis Hoplophthiracarus Jacot, 1938 synonym of Hoplophthiracarus illinoisensis (Ewing, 1909)

pandanensis Notophthiracarus Niedbała et Starý, 2014

Notophthiracarus pandanensis: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) pandanensis: Subías 2022 .

Afrotropic: Madagascar (Niedbała 2017 a, Niedbała & Starý 2014b, Subías 2022).

Afrotropical, Madagascar, endemic pandus Phthiracarus Niedbała, 2004

Phthiracarus pandus: Niedbała & Liu 2018 .

Phthiracarus (Archiphthiracarus) pandus: Subías 2022 .

Neotropic: Jamaica ( Niedbała 2004a, Niedbała & Starý 2015e, Subías 2022).

Neotropical, Jamaica, endemic pannonicus Phthiracarus Willmann , in: Franz 1954 synonym of Phthiracarus longulus (C.L. Koch, 1841)

pantotrema Apoplophora ( Berlese, 1913)

Mesoplophora pantotrema Berlese, 1913

Mesoplophora pantotrema: Hammen 1959 , Hammer 1979, Castagnoli & Pegazzano 1985.

Apoplophora pantotrema: Niedbała 1984a, 1993a , 1998b, 2000, 2001c, 2004c, 2006d, 2007a, 2011b, 2012b, Niedbała & Corpuz-Raros 1998, Wang et al. 2002, Niedbała et al. 2006, Niedbała & Penttinen 2007, Liu et al. 2009a, Chen et al. 2010, Liu 2016a, 2021b, Niedbała & Liu 2018, Niedbała & Ermilov 2018a, 2022, Subías 2022.

Mesoplophora discreta Berlese, 1913 syn. by Hammen 1959.

Mesoplophora discreta Berlese, 1913 : Castagnoli & Pegazzano 1985.

Mesoplophora rostrorugosa Hammer, 1979 syn. by Niedbała 1984a.

Apoplophora remota Aoki, 1980 syn. by Niedbała 1984a.

Mesoplophora indica Mahunka, 1985 syn. by Niedbała 2000.

Mesoplophora striata Mahunka, 1985 syn. by Niedbała 2000.

Apoplophora lineata Mahunka, 1987 syn. by Niedbała 2000.

Apoplophora lineata: Mahunka 1991b .

Apoplophora ornatissima Mahunka, 1988 syn. by Niedbała 2000.

Apoplophora jaccoudi Mahunka, 1991 syn. by Niedbała 2000.

Palaearctic: Japan, North India (Assam, Arunachal Pradesh), Nepal, China (Beijing, Jilin, Heilongjiang). Orient: China (Fujian, Hainan, Taiwan, Macau Peninsula), Andaman Islands, South India, Sri Lanka, Thailand, Southern and Northern Vietnam, Malaysia ( Sarawak, Pahang, West Malaysia), Indonesia (Bali, Java, Sumatra, Sulawesi, Lombok), Phillipines. Australasia: Papua New Guinea, Australia (northern and western parts, Tasmania), Solomon Islands, Fiji ( Aoki 1965, 1980, Balogh & Balogh 2002, Berlese 1913, Castagnoli & Pegazzano 1985, Chen et al. 2010, Ermilov et al. 2012, Hammen 1959, Hammer 1979, 1982, Liu 2016a, 2021b, Liu et al. 2009a, Mahunka 1985d, 1987f, 1988d, 1991b, c, Mahunka & Mahunka-Papp 1995, Niedbała, 1981, 1984a, 1998b, 1993a, 2000, 2001c, 2004c, 2006d, 2007a, 2008b, 2011a, b, 2012a,

b, 2014b, 2017b, Niedbała & Corpuz-Raros 1998, Niedbała & Ermilov 2013a, b, 2014b, 2017a, 2018a, b, 2020, 2021, 2022, Niedbała et al. 2006, Niedbała & Penttinen 2007, Niedbała & Starý 2015e, Ryabinin et al. 2018, Subías 2022, Vu 2013, Wang et al. 2002,).

Oriental (broad) introduced to eastern and southern Palaearctic, northern and western Pacific Islands. Pantropical ?

papillatus Atropacarus Gil-Martin, Subías et Arillo, 1992 synonym of Atropacarus (Atropacarus) platakisi ( Mahunka, 1979)

papillosus Austrophthiracarus (Parry, 1979)

Phthiracarus papillosus Parry, 1979

Phthiracarus papillosus: Niedbała 1986b, 1992 .

Archiphthiracarus papillosus: Balogh & Mahunka 1983 .

Phthiracarus (Archiphthiracarus) papillosus: Subías 2004 .

Austrophthiracarus papillosus: Niedbała 2011b, 2012b , Liu & Zhang 2016 a, Niedbała & Liu 2018. Hoplophthiracarus (Notophthiracarus) papillosus: Subías 2022 .

Palaearctic: Austria ( Balogh & Mahunka 1983, Niedbała 1986b, 1992, 2011b, 2012 b, Parry 1979 b, Schatz 1983, Subías 2022).

Western Palaearctic, Austria, endemic paraafricanus Euphthiracarus (Pocsia) Niedbała et Starý, 2014

Euphthiracarus (Pocsia) paraafricanus: Niedbała & Liu 2018 , Subías 2022.

Afrotropic: Tanzania (Niedbała 2014a, 2017c, Niedbała & Starý 2014c, Subías 2022).

Afrotropical, Tanzania, endemic paraajaluela Oribotritia (Oribotritia) Niedbała et Illig, 2007 bad spelling of Oribotritia (Oribotritia) paraalajuela Niedbała et Illig, 2007

paraalajuela Oribotritia (Oribotritia) Niedbała et Illig, 2007

Oribotritia paraalajuela Niedbała et Illig, 2007

Oribotritia paraajaluela: Liu 2015 a, Niedbała & Liu 2018 .

Oribotritia (Oribotritia) paraajaluela (lapsus linguae): Subías 2022.

Neotropic: Ecuador ( Niedbała & Illig 2007b, Niedbała & Roszkowska 2017, Subías 2022).

Neotropical, Ecuador, endemic.

Note. The name of this species refers to the similarity to O. alajuela Niedbała, 2003 . It was mistakenly written as O. paraajaluela at the description of species and at the legend of figures, but correctly used in abstract.

paraallocotos Arphthicarus Niedbała, 2006

Arphthicarus paraallocotos: Niedbała 2008e , Niedbała & Liu 2018.

Hoplophthiracarus (Notophthiracarus) paraallocotos: Subías 2022 .

Neotropic: Costa Rica ( Niedbała 2006e, 2008e, Subías 2022).

Neotropical, Costa Rica, endemic paraandinus Austrophthiracarus (Subías, 2018)

nov. nom for Austrophthiracarus baloghi Niedbała, 2003 : Subías 2004 (updated in 2018). Austrophthiracarus paraandinus: Niedbała & Liu 2018 .

Notophthiracarus (Protophthiracarus) paraandinus Subías, 2018

Austrophthiracarus baloghi Niedbała, 2003 nom. praeoc. by Niedbała 1987.

Austrophthiracarus baloghi: Niedbała 2004a .

Hoplophthiracarus (Protophthiracarus) paraandinus: Subías 2022 .

Neotropic: Chile ( Niedbała 2003b, 2004a, Subías 2022, 2009).

Neotropical, Chile, endemic paraaokii Oribotritia (Oribotritia) Niedbała, 2000

Oribotritia paraaokii Niedbała, 2000

Oribotritia paraaokii: Subías 2004 , Liu 2015 a, Niedbała & Liu 2018.

Oribotritia (Oribotritia) paraaokii: Subías 2022 .

Orient: Northern Vietnam ( Niedbała 2000, Subías 2022).