Indotritia lebronneci, Jacot, 1934

Niedbała, Wojciech & Liu, Dong, 2023, Systematic, synonymic and biogeographical list of ptyctimous mites (Acari, Oribatida) in the world (1799 – 2022), Zootaxa 5265 (1), pp. 1-442 : 126-134

Pantropical, eastern part lebronneci crassiori Indotritia Jacot, 1934 = Austrotritia lebronneci (Jacot, 1934)

lebronneci flagelloides Indotritia Jacot, 1934 = Austrotritia lebronneci (Jacot, 1934)

lebronneci tahitiana Indotritia Jacot, 1934 = Austrotritia lebronneci (Jacot, 1935)

leei Notophthiracarus Niedbała, 1987

Notophthiracarus lee Niedbała, 1987 (lapsus linguae)

Notophthiracarus lee (lapsus linguae): Niedbała 1992, 2000, Liu & Zhang 2014d, Niedbała & SzywilewskaSzczykutowicz 2017.

Notophthiracarus leei: Niedbała & Liu 2018 , Niedbała et al. 2021.

Notophthiracarus (Notophthiracarus) leei: Subías 2004 .

Hoplophorella (Hoplophorella) leei: Subías 2022 .

Australasia: Australia (western and southern parts) ( Balogh & Balogh 2002, Niedbała 1987, 1992, 2000, 2012a, 2017b, Niedbała et al. 2021, Niedbała & Szywilewska-Szczykutowicz 2017, Subías 2022).

Australasian, western and southern parts of Australia leliehoekensis Phthiracarus Niedbała, 2006

Phthiracarus leliehoekensis: Niedbała & Liu 2018 .

Phthiracarus (Phthiracarus) leliehoekensis: Subías 2022 .

South Africa: Free State ( Niedbała 2006c, Subías 2022).

South African, endemic lemurius Atropacarus (Hoplophorella) Mahunka, 1993

Hoplophorella lemuria Mahunka, 1993

Atropacarus (Hoplophorella) lemurius: Niedbała 2001a, 2004 d , Niedbała & Liu 2018.

Hoplophorella (Hoplophorella) lemuria: Subías 2004 .

Hoplophorella (Kakophthiracarus) lemuria: Subías 2022 .

Afrotropic: Madagascar ( Balogh & Balogh 2002, Mahunka 1993c, 2002, Niedbała 2001a, 2004d, 2017a, Subías 2022).

Afrotropical, Madagascar, endemic lentula Tritia sensu Berlese, 1887 species inquirenda, incertae sedis

Tritia lentula: Castagnoli & Pegazzano 1985 , Niedbała 2011b, Niedbała & Liu 2018.

Acrotritia lentula: Subías 2004 .

Palaearctic: Italy ( Berlese 1887b)

lentula var. hyeroglyphica (K.) Tritia Berlese, 1916 synonym of Acrotritia ardua (C.L. Koch, 1841)

lentulus Phthiracarus (C.L. Koch, 1841)

Hoplophora lentula C.L. Koch, 1841

Tritia lentula: Berlese 1887b .

Phthiracarus lentulus: Jacot 1936 , Feider & Călugăr 1970, Krivolutsky 1975, Balogh & Mahunka 1980 a, 1983, Kamill 1981, Niedbała 1986b, 1992, 2000, 2001a, 2002a, 2007c, 2008f, 2011b, 2012b, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008, Mahunka & Mahunka-Papp 2004, Ni & Liu 2014, Niedbała & Liu 2018, Liu 2021b.

Phthiracarus (Phthiracarus) lentulus: Subías 2022 .

Phthiracarus rotundus Berlese, 1923 nom. praeoc. by Ewing, 1908: Subías 2022.

Phthiracarus angolensis Mahunka, 1985 syn. by Niedbała 2001a.

Phthiracarus angolensis: Niedbała 1992 .

? Phthiracarus italicus sensu Feider et Suciu, 1957

Nearctic: Canada (Ontario, Quebec, Nova Scotia ), USA (Oregon, Texas, Illinois, Michigan, Indiana, Kentucky, Pennsylvania, New York, West Virginia). Palaearctic: Spain, Norway, Finland, Denmark, Germany, Luxembourg, Switzerland, Austria, Italy, Poland, Czech Republic, Slovakia, Hungary, Romania, Bulgaria, Slovenia, Greece, Turkey, Ukraine (western part), Georgia, Russia (Central European Territory, Far East), Algeria, Iran, Turkmenistan, China (Neimenggu, Xinjiang), North Korea. Afrotropic: Angola, Madagascar (central and eastern parts). Orient: China (Sichuan, Hubei, Chongqing, Fujian, Guizhou, Hainan) ( Balogh & Balogh 2002, Balogh & Mahunka 1980 a, 1983, Berlese 1887 b, Castagnoli & Pegazzano 1985, Jacot 1936, Kamill 1981, C.L. Koch 1841, Liu 2021b, Mahunka 1985a, Mahunka et al. 2013, Mahunka & Mahunka-Papp 1999, 2003, 2004, Mahunka & Paoletti 1984, Niedbała 1986b, 1992, 2000, 2001a, 2002a,

2007c, 2008f, 2011b, 2012b, 2015b, 2017a, Niedbała & Ermilov 2018b, Olszanowski et al. 1996, Schatz 1983, Seniczak et al. 2021, Subías 2022, Weigmann et al. 2015).

Holarctic, Panpalaearctic introduced to Northern Oriental and Afrotropic ( Angola, Madagascar)

Notes. Probably P. lentulus (C.L.Koch, 1841) and Hoploderma italicum Oudemans, 1900 are congeneric. However, it is not possible to establish such a synonymy on the basis of description of Oudemans (1900). Subías (2022) considered this species as congeneric with P. incertus Niedbała, 1983 (= P. conformis Sergenko, 1987 ). However, both species have slightly different shape of trichobothria, and mutual distance between rostral setae is shorter and notogastral setae are longer in P. lentulus .

Subías (2022) considered Phthiracarus rotundus Berlese, 1923 as congeneric with this species. See note of P. rotundus .

leonilae Plonaphacarus Niedbała, 2017

Plonaphacarus leonilae: Niedbała & Ermilov 2017 a, Niedbała & Liu 2018 .

Hoplophthiracarus (Hoplophthiracarus) leonilae: Subías 2022 .

Orient: Philippines ( Niedbała & Ermilov 2017a, Subías 2022).

Oriental, Philippines, endemic lepteces Oribotritia (Oribotritia) Niedbała, Corpuz-Raros et Gruezo, 2006

Oribotritia lepteces Niedbała, Corpuz-Raros et Gruezo, 2006

Oribotritia lepteces: Liu 2015a, 2016b , Niedbała & Liu 2018.

Oribotritia (Oribotritia) lepteces: Subías 2022 .

Orient: Myanmar, Philippines, Malaysia. Australasia: Australia (New South Wales) ( Niedbała 2012 a, 2014b, Niedbała & Ermilov 2018b, Niedbała et al. 2006, Niedbała & Starý 2015e, Subías 2022).

Oriental, Philippinian and Australasian levidensus Atropacarus (Hoplophorella) Niedbała et Starý, 2022

Afrotropic: Malawi ( Niedbała & Starý 2022)

Afrotropical, Malawi, endemic leviseta Mesoplophora (Parplophora) Hammer, 1979

Mesoplophora leviseta Hammer, 1979

Mesoplophora (Parplophora) leviseta: Niedbała 1985a, 1998b , 2000, 2001a, 2010a, Liu & Chen 2010c, Liu 2016 a, Niedbała & Liu 2018, Subías 2022.

Mesoplophora gibba Mahunka, 1988 syn. by Niedbała 2000.

Afrotropic: Seychelles. Orient: Indonesia (Sumatra, Java). Australasia: Fiji, Tahiti ( Balogh & Balogh 2002, Hammer 1979, Mahunka 1988b, Niedbała 1985a, 1998b, 2000, 2001a, 2010a, 2012a, 2017a, Niedbała & Ermilov 2020, Niedbała & Starý 2015e, Subías 2022).

Pantropical lewisensis Notophthiracarus Niedbała, 2012

Notophthiracarus lewisensis: Liu & Zhang 2014 d, Niedbała & Liu 2018 .

Hoplophthiracarus (Steganacarellus) lewisensis: Subías 2022 .

Australasia: Australia (Queensland) ( Niedbała 2012a, Subías 2022).

Australasian, Australia, endemic liaoi Phthiracarus Niedbała, 2021

Phthiracarus (Archiphthiracarus) liaoi: Subías 2022 .

Orient: China (Taiwan) ( Niedbała & Ermilov 2021).

Oriental, China (Taiwan), endemic licnophora var. obsoletior Phthiracarus (Hoplophorella) Berlese, 1923 synonym of Atropacarus (Hoplophorella) hamatus (Ewing, 1909)

licnophorum Hoploderma Berlese, 1913 synonym of Atropacarus (Hoplophorella) cucullatus (Ewing, 1909) lienhardi Hoplophorella Mahunka, 1987 synonym of Atropacarus (Hoplophorella) vitrinus ( Berlese, 1913)

Note. Subías (2022) incorrectly synonymized this species with Atropacarus (Hoplophorella) andrei ( Balogh, 1958) .

lienhardi Notophthiracarus Mahunka, 1996

Notophthiracarus lienhardi: Niedbała 2000, 2007 a, Niedbała & Liu 2018 .

Notophthiracarus (Besuchetacarus) lienhardi: Subías 2004 .

Hoplophthiracarus (Besuchetacarus) lienhardi: Subías 2022 .

Orient: Malaysia (Perak, Sarawak, Pahang), Vietnam (Hanoi), China (Hainan) ( Balogh & Balogh 2002, Mahunka 1996a, Niedbała 2000, 2007a, Niedbała & Ermilov 2018b, 2020, Niedbała & Starý 2015e, Subías 2022).

Oriental, Malayan lienhardi Sabahtritia Mahunka, 1995

Sabahtritia lienhardi: Mahunka 1995b , Niedbała 2000, Liu & Chen 2009, Niedbała & Liu 2018, Subías 2022.

Orient: Brunei, Malaysia ( Sarawak) ( Mahunka 1995b, Niedbała 2000, 2008b, Subías 2022).

Oriental, Malayan ligneus Phthiracarus Willmann, 1932 synonym of Phthiracarus ferrugineus (C.L. Koch, 1841)

ligulifera Hoplophorella Mahunka, 1987 synonym of Atropacarus (Hoplophorella) hamatus (Ewing, 1909) lineata Apoplophora Mahunka, 1987 synonym of Apoplophora pantotrema ( Berlese, 1913)

lineatus Notophthiracarus Niedbała et Starý, 2015

Notophthiracarus lineatus: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) lineatus: Subías 2022 .

Afrotropic: Madagascar ( Niedbała & Starý 2015 d, Subías 2022).

Afrotropical, Madagascar, endemic lineolatus Notophthiracarus Mahunka, 1993

Notophthiracarus lineolatus: Niedbała 2001 a, Niedbała & Liu 2018 .

Notophthiracarus (Notophthiracarus) lineolatus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) lineolatus: Subías 2022 .

Afrotropic: Tanzania ( Balogh & Balogh 2002, Mahunka 1993d, Niedbała 2001a, 2017c, Subías 2022). Afrotropical, Tanzania, endemic lionsi Notophthiracarus Niedbała, 1987

Notophthiracarus lionsi: Niedbała 1992, 2000 , Liu & Zhang 2014 d, Niedbała & Liu 2018. Notophthiracarus (Notophthiracarus) lionsi: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) lionsi: Subías 2022 .

Australasia: Australia (Queensland) ( Balogh & Balogh 2002, Niedbała 1987, 1992, 2000, 2012a, Subías 2022).

Australasian, Australia, endemic liparus Phthiracarus Niedbała, 1984

Phthiracarus liparus: Niedbała 1986b, 1992 , 2011b, 2012b, Niedbała & Liu 2018.

Phthiracarus (Archiphthiracarus) liparus: Subías 2022 .

Palaearctic: Canary Islands, Algeria, Turkey ( Niedbała 1984f, 1986b, 1992, 2011b, 2012b, Subías 2022). Western Palaearctic, distributed in Macaronesia, Maghrebian and Anatolian areas, disjunctive liratus Notophthiracarus Niedbała et Starý, 2015

Notophthiracarus liratus: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) liratus: Subías 2022 .

Afrotropic: Madagascar (Niedbała 2017a, Niedbała & Starý 2015 d, Subías 2022).

Afrotropical, Madagascar, endemic lividus Hoplophthiracarus Niedbała, 2000

Hoplophthiracarus lividus: Liu & Zhang 2014c , Niedbała & Liu 2018.

Rhacaplacarus (Rhacaplacarus) lividus: Subías 2004 .

Phthiracarus (Archiphthiracarus) lividus: Subías 2022 .

Australasia: New Zealand ( Niedbała 2000, 2012a, Subías 2022).

Australasian, New Zealand, endemic loebli Plonaphacarus (Mahunka, 1985)

Hoplophthiracarus loebli Mahunka, 1985

Hoplophthiracarus loebli: Niedbała 1992 .

Plonaphacarus loebli: Niedbała 1994a, 2000 , 2007a, Liu et al. 2011b, Niedbała & Liu 2018. Rhacaplacarus (Rhacaplacarus) loebli: Subías 2004 .

Hoplophthiracarus (Hoplophthiracarus) loebli: Subías 2022 .

Orient: India (Kerala), Indonesia (Sumatra, Sulawesi) ( Balogh & Balogh 2002, Mahunka 1985 d, Niedbała 1992, 1994a, 2000, 2007a, Niedbała & Ermilov 2020, Subías 2022).

Oriental longicarinatus Atropacarus (Hoplophorella) Liu, 2021

Atropacarus (Hoplophorella) longicarinatus: Liu 2021b .

Hoplophorella (Hoplophorella) longicarinata: Subías 2022 .

Orient: China (Guangxi) (Liu 2021a, b).

Oriental, China, endemic longipilosus Phthiracarus Niedbała, 2021

Phthiracarus (Archiphthiracarus) longipilosus: Subías 2022 .

South Africa: Eastern Cape (Niedbała et al. 2021).

South African, endemic longicarinatus Plonaphacarus Niedbała, 2008

Plonaphacarus longicarinatus: Liu et al. 2011b , Niedbała & Liu 2018.

Hoplophthiracarus (Hoplophthiracarus) longicarinatus: Subías 2022 .

Orient: Malaysia ( Sarawak) (Liu et al. 2013, Niedbała 2008b, Subías 2022).

Oriental, Malaysia, endemic longipila Synichotritia Niedbała, 2002

Synichotritia longipila: Niedbała & Liu 2018 , Subías 2022.

Nearctic: USA (Mississippi, Tennessee, Illinois) ( Niedbała 2002a, Niedbała & Starý 2015e, Subías 2022). Nearctic, disjunctive longipilosus Steganacarus (Rhacaplacarus) Niedbała, 2004

Steganacarus (Rhacaplacarus) longipilosus: Niedbała & Liu 2018 .

Hoplophorella (Rhacaplacarus) longipilosa: Subías 2022 .

Neotropic: Brazil ( Niedbała 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Neotropical, Brazil, endemic longirostralis Euphthiracarus (Euphthiracarus) Walker, 1965

Euphthiracarus longirostralis Walker, 1965

Euphthiracarus longirostralis: Niedbała 2002a, 2007c , Niedbała & Liu 2018.

Euphthiracarus (Euphthiracarus) longirostralis: Subías 2022 .

Nearctic: Canada (British Columbia), USA (Oregon, California) ( Marshall et al. 1987, Niedbała 2002a, 2007c, Niedbała & Starý 2015e, Subías 2022, Walker 1965).

Nearctic, western part longisensillus Phrathicarus Liu et Zhang, 2013

Phrathicarus longisensillus: Niedbała & Liu 2018 .

Atropacarus (Atropacarus) longisensillus: Subías 2022 .

Australasia: New Zealand ( Liu & Zhang 2013b, Subías 2022).

Australasian, New Zealand, endemic longiseta Mesoplophora (Mesoplophora) Mahunka, 1982

Mesoplophora longiseta Mahunka, 1982

Mesoplophora longiseta: Niedbała 2003a .

Mesoplophora (Mesoplophora) longiseta: Niedbała 1985a, 2004 a , Niedbała & Liu 2018, Subías 2022. Neotropic: Costa Rica ( Balogh & Balogh 1988, 2002, Mahunka 1982e, Niedbała 1985a, 2003a, 2004a, Subías 2022).

Neotropical, Costa Rica, endemic

Note. After supposition of Niedbała (1985a), Subías (2004) proposed this species as congeneric with Mesoplophora cubana Călugăr et Vasiliu, 1977 . However, these two species differ by the presence of notogastral setae smooth ( M. cubana ) and ciliate ( M. longiseta ). Subías (2022) retreated from this opinion.

longisetosa Mesoplophora (Mesoplophora) Călugăr et Vasiliu, 1977

Mesoplophora (Mesoplophora) longisetosa: Niedbała 1985a, 2004a , Liu & Chen 2010c, Liu 2016 a, Niedbała & Liu 2018, Subías 2022.

Neotropic: Antilles ( Cuba) ( Balogh & Balogh 1988, 2002, Călugăr & Vasiliu 1977, Niedbała 1985a, 2004a, Subías 2022).

Neotropical, Cuba, endemic longisetosus Austrophthiracarus Liu et Chen, 2014

Austrophthiracarus longisetosus: Liu & Zhang 2016 a, Niedbała & Liu 2018 , Liu 2021b. Hoplophthiracarus (Calyptophthiracarus) longisetosus: Subías 2022 .

Orient: China (Hubei) (Liu 2021b, Liu & Chen 2014b, Niedbała 2015b, Subías 2022).

Oriental, China, endemic longisetosus Austrophthiracarus Niedbała et Starý 2015 junior homonym of Austrophthiracarus longisetosus Liu et Chen, 2014 changed on

Austrophthiracarus longisetus Niedbała et Starý, 2015

longisetosus Steganacarus Moraza, 1984 synonym of Steganacarus (Steganacarus) herculeanus Willmann, 1953 longisetosus Plonaphacarus Niedbała, 2019

Hoplophorella (Rhacaplacarus) longisetosa: Subías 2022 .

Afrotropic: Ivory Coast ( Niedbała & Ermilov 2019a).

Afrotropical, Ivory Coast, endemic longisetus Austrophthiracarus Niedbała et Starý 2015

Austrophthiracarus longisetus: Niedbała & Starý 2015e , Niedbała & Liu 2018.

Austrophthiracarus longisetosus: Niedbała & Starý 2015b .

Phthiracarus (Neophthiracarus) longisetus: Subías 2022 .

Austrophthiracarus niedbalai Liu & Zhang, 2016 syn. by Niedbała & Liu, 2018.

Neotropic: Bolivia ( Niedbała & Starý 2015b, Subías 2022).

Neotropical, Bolivia, endemic longisetus Euphthiracarus (Euphthiracarus) Liu, Wu et Chen, 2011

Euphthiracarus longisetus Liu, Wu et Chen, 2011

Euphthiracarus longisetus: Liu et al. 2011 d, 2012 , Niedbała & Liu 2018, Liu 2021b.

Euphthiracarus (Euphthiracarus) longisetus: Subías 2022 .

Orient: China (Hunan) (Liu 2021b, Liu et al. 2011d, 2021, 2013, Subías 2022).

Oriental, China, endemic longisetus Notophthiracarus Niedbała, 2000

Notophthiracarus longisetus: Liu & Zhang 2014 d, Niedbała & Liu 2018 .

Notophthiracarus (Notophthiracarus) longisetus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) longisetus: Subías 2022 .

Australasia: Australia (Queensland) ( Niedbała 2000, 2012a, Subías 2022).

Australasian, Australia, endemic longqiensis Hoplophthiracarus Liu, 2015

Hoplophthiracarus longqiensis: Niedbała & Liu 2018 , Liu 2021b.

Hoplophorella (Rhacaplacarus) longqiensis: Subías 2022 .

Orient: China (Fujian) (Liu 2015b, 2021b).

Oriental, China, endemic longulus Euphthiracarus (Euphthiracarus) Liu, Wu et Chen, 2011

Euphthiracarus longulus Liu, Wu et Chen, 2011

Euphthiracarus longulus: Liu et al. 2011 d, 2012 , Niedbała & Liu 2018, Liu 2021b.

Euphthiracarus (Euphthiracarus) longulus: Subías 2022 .

Orient: China (Fujian) (Liu 2021b, Liu et al. 2011 d, 2012, 2013, Subías 2022).

Oriental, China, endemic longulus Phthiracarus (C.L. Koch, 1841)

Hoplophora longula C.L. Koch, 1841

Hoplophora longula: Oudemans 1937 .

Phthiracarus longulus: Kamill 1981 , Niedbała 1986b, 1992, 2000, 2001b, 2002a, 2008e, f, 2010b, 2011b, 2012b, Olszanowski et al. 1996, Niedbała & Olszanowski 1997, 2004, 2008, Mahunka & Mahunka-Papp

2004, Weigmann 2006, Ni & Liu 2014, Niedbała & Liu 2018, Liu 2021b, Niedbała et al. 2021, Niedbała & Błoszyk 2022.

Phthiracarus (Phthiracarus) longulus: Mahunka 1990b , Subías 2022.

Phthiracarus apiculatus: Niedbała 1986b .

Phthiracarus apiculatus Jacot, 1939 syn. by Niedbała 2001b.

Phthiracarus apiculatus: Marshall et al. 1987 , Niedbała 1992.

Phthiracarus ferrugineus (C.L. Koch) sensu Jacot, 1936 (in part).

Phthiracarus flexisetosus Parry, 1979 syn. by Niedbała 1986b.

Phthiracarus flexisetosus: Balogh & Mahunka 1983 , Mahunka & Mahunka-Papp 2004.

Phthiracarus hortobagyensis Mahunka, 1982 syn. by Niedbała 1986b.

Phthiracarus hortobagyensis: Mahunka & Mahunka-Papp 2004 .

Phthiracarus montium Jacot, 1939 syn. by Niedbała 2001b.

Phthiracarus montium: Marshall et al. 1987 .

Phthiracarus mundus Niedbała, 1983 syn. by Niedbała 1986b.

Phthiracarus pannonicus Willmann in: Franz 1954 syn. by Niedbała 1986b.

Phthiracarus pannonicus Willmann in: Franz 1954: Subías 2012

Phthiracarus prior Jacot, 1933 syn. by Niedbała 2001b.

Phthiracarus prior: Niedbała 1986b, 1992 , 1994a, Marshall et al. 1987.

Phthiracarus setosellus Jacot, 1929 syn. by Niedbała 1986b.

Phthiracarus setosellus: Jacot 1930 , Marshall et al. 1987.

Phthiracarus tardus Forsslund, 1943 syn. by Niedbała 1986b.

Phthiracarus tardus: Forsslund 1945, 1956 , Parry 1979a, Balogh & Mahunka 1980 a, 1983.

Nearctic: Canada (Ontario, British Columbia), USA (California, North Carolina, New York, Utah, New Mexico, Michigan, Kentucky, Mississippi, Alabama, Florida, Pennsylvania, Tennessee, Utah). Neotropic: Uruguay. Palaearctic: England, Spain, France, the Netherlands, Norway, Sweden, Finland, Denmark, Germany, Luxembourg, Switzerland, Italy, Austria, Poland, Czech Republic, Slovakia, Hungary, Croatia, Greece, Romania, Estonia, Latvia, Lithuania, Belarus, Ukraine (western and eastern part and Crimea), Azerbaijan, Georgia, Russia (Northern, Central and South European Territories, Western and Eastern Siberia, Far East), Canary Islands, Algeria, Turkey, Kazakhstan, Uzbekistan, Kyrgyzstan, China (Tibet, Xinjiang) ( Balogh & Mahunka 1980 a, 1983, Beck et al. 2014, Forsslund 1956, Franz 1954, Jacot 1929, 1930, 1933, 1937, 1939, Kamill 1981, Liu 2021b, Mahunka 1982b, 1990 b, Mahunka & Mahunka-Papp 1999, 2003, 2004, 2008f, Mahunka et al. 2013, Marshall et al. 1987, Niedbała 1983f, 1986b, 1992, 1994a, 2000, 2001b, 2002a, 2008e, f, 2010b, 2011b, 2012b, 2015b, 2018, Niedbała & Błoszyk 2022, Niedbała et al. 2014, 2021, Niedbała & Starý 2015e, Norton & Macnamara 1976, Olszanowski et al. 1996, Oudemans 1937, Parry 1979 a, Schatz 1983, Schmelzle et al. 2010, Seniczak et al. 2021, Subías 2022, Weigmann et al. 2015).

Holarctic, Panpalaearctic, probably introduced to Uruguay

Notes. Subías (2022) considered Hoploderma boreale Trägårdh, 1910 , Phthiracarus crenophilus Willmann, 1951 and Phthiracarus malagensis Niedbała, 1986 as congeneric with this species. However, it is not possible to establish such a synonymy with Hoploderma boreale on the basis of description and figures of Trägårdh (1910).

Notogastral setae are shorter (c 1 <c 1 - d 1) in P. crenophilus versus c 1> c 1 - d 1 in P.

longulus and presence of four setae on femora of legs I versus three setae in P. longulus .

P. malagensis has rounded distal end of trichobothria (versus pointed end in P. longulus ), longer notogastral setae and chaetome of legs complete (versus chaetome incomplete).

longus Austrophthiracarus Niedbała, 2017

Austrophthiracarus longus: Niedbała & Liu 2018 .

Hoplophthiracarus (Notophthiracarus) longus: Subías 2022 .

Australasia: Australia (Queensland) (Niedbała 2017b, Subías 2022).

Australasian, Australia, endemic loricatum Trombidium Rathke, 1799 species inquirenda, incertae sedis

Trombidium loricatum: Niedbała 2011b , Niedbała & Liu 2018.

Hoplophora loricatum: Oudemans 1937

Acrotritia loricata: Subías 2004 .

Oribotritia loricata: Willmann 1931 , Hammen 1952.

Palaearctic, Germany ( Rathke 1799, Oudemans 1937)

Note. Subías (2022) considered this species as congeneric with A. ardua (C.L. Koch, 1841) .

lucida Hoplophora C.L. Koch, 1841 species inquirenda

Hoplophora lucida: Sellnick 1928 , Willmann 1931, Oudemans 1937, Hammen 1952, Feider & Suciu 1967, Niedbała 2011b, Niedbała & Liu 2018.

Archiphthiracarus lucidus: Balogh & Mahunka 1980 a, 1983 .

Phthiracarus (Phthiracarus) piger: Subías 2004 .

Phthiracarus (Archiphthiracarus) piger: Subías 2022 .

Palaearctic: Germany ( Balogh & Mahunka 1983, Niedbała 1992).

Note. The type specimen does not exist but the name was used by many authors ( Sellnick 1928, Willmann 1931, Oudemans 1937, Hammen 1952, Feider & Suciu 1967, Krivolutsky 1975, Balogh & Mahunka 1980 a, 1983, Berg et al. 1990, Subías 2004).

Note. Subías (2022) considered Acarus piger Scopoli, 1763 as congeneric with this species. It is not possible to synonymize these two species inquirenda.

lucida Rhysotritia Niedbała, 1998 synonym of Acrotritia curticephala ( Jacot, 1938)

lucidus Steganacarus Sergienko, 1994 synonym of Steganacarus (Tropacarus) carinatus (C.L. Koch, 1841) (?) Note. Subías (2022) considered Steganacarus (Tropacarus) malatestai Avanzati, Baratti et Bernini, 1995 as congeneric with this species. See at S. (T.) malatestai .

lunatus Austrophthiracarus Niedbała, 2017

Austrophthiracarus lunatus: Niedbała & Szywilewska-Szczykutowicz 2017 , Niedbała & Liu 2018, Niedbała et al. 2021.

Hoplophthiracarus (Protophthiracarus) lunatus: Subías 2022 .

Australasia: Australia (Victoria) (Niedbała et al. 2021, Niedbała & Szywilewska-Szczykutowicz 2017, Subías 2022).

Australasian, Australia, endemic lunatus Notophthiracarus Niedbała, 2004

Notophthiracarus lunatus: Niedbała & Liu 2018 .

Hoplophthiracarus (Steganacarellus) lunatus: Subías 2022 .

Neotropic: Brazil ( Niedbała 2004 a, Oliveira 2004, Oliveira et al. 2017, Subías 2022).

Neotropical, Brazil, endemic luridus Phthiracarus (Ewing, 1909)

Hoploderma lurida Ewing, 1909

Hoploderma lurida: Niedbała 1986b, 1992 .

Ginglymacarus lurida: Ewing 1917b .

Phthiracarus luridus: Jacot 1938 a , Niedbała 2002 a, Niedbała & Liu 2018.

Phthiracarus (Phthiracarus) luridus: Subías 2004 .

Phthiracarus (Archiphthiracarus) luridus: Subías 2022 .

Nearctic: Canada (British Columbia, Quebec), USA (Illinois, Oregon, California) ( Ewing 1909c, 1917b, Jacot 1938, Marshall et al. 1987, Niedbała 1986b, 1992, 2002a, Subías 2022).

Nearctic, northern and western parts lutulentus Phthiracarus Sellnick, 1920 synonym of Atropacarus (Atropacarus) csiszarae ( Balogh et Mahunka, 1979)

luxiensis Plonaphacarus Liu et Huang, 2021

Hoplophthiracarus (Steganacarellus) luxiensis: Subías 2022 .

Orient: China (Hunan) ( Liu & Huang 2021a).

Oriental, China, endemic machadoi Phthiracarus Balogh, 1958 nomen nudum

Phthiracarus machadoi: Niedbała 2001 a, Niedbała & Liu 2018 .

Phthiracarus (Phthiracarus) machadoi: Subías 2022 .

Afrotropic: Angola ( Balogh 1958, Niedbała 2001a, Subías 2022).

Afrotropical, Angola, endemic machadoi Plonaphacarus ( Balogh, 1958)

Steganacarus machadoi Balogh, 1958

Hoplophthiracarus machadoi: Mahunka 1984b .

Plonaphacarus machadoi: Niedbała 1986b, 1992 , 2001a, Liu et al. 2011b, Niedbała & Liu 2018, Niedbała & Ermilov 2022c.

Hoplophthiracarus (Plonaphacarus) machadoi: Mahunka 1992 .

Rhacaplacarus (Rhacaplacarus) machadoi: Subías 2004 .

Hoplophorella (Rhacaplacarus) machadoi: Subías 2022 .

Rhacaplacarus machadoi: Mahunka & Mahunka-Papp 2009a .

Afrotropic: Ivory Coast, Cameroon, Ethiopia, Democratic Republic Congo, Uganda, Kenya, Rwanda, Tanzania, Angola, Zambia, Malawi, Madagascar ( Balogh 1958, Mahunka 1984b, 1992, Mahunka & Mahunka-Papp 2009a, Niedbała 1986b, 1992, 2001a, 2008c, 2017a, c, Niedbała & Ermilov 2011, 2013, 2014a, 2022c, Niedbała & Starý 2022, Subías 2022).

Afrotropical, central and eastern parts macrorostralis Euphthiracarus (Euphthiracarus) Liu, Wu et Chen, 2012

Euphthiracarus macrorostralis Liu, Wu et Chen, 2012

Euphthiracarus macrorostralis: Liu et al. 2012 , Niedbała & Liu 2018, Liu 2021b.

Euphthiracarus (Euphthiracarus) macrorostralis: Subías 2022 .

Orient: China (Sichuan) (Liu 2021b, Liu et al. 2012, 2013, Niedbała 2015b, Subías 2022).

Oriental, China, endemic macrosculpturatus Steganacarus (Steganacarus) Mahunka et Mahunka Papp, 1999 synonym of Steganacarus (Steganacarus) balearicus Pérez-Íñigo, 1969

maculatus Notophthiracarus (Trägårdh, 1931)

Phthiracarus maculatus Trägårdh, 1931

Phthiracarus maculatus: Niedbała 1986b, 1992 .

Notophthiracarus maculatus: Niedbała 1994a, 2004 a , Niedbała & Liu 2018.

Notophthiracarus (Notophthiracarus) maculatus: Subías 2004 .

Hoplophthiracarus (Notophthiracarus) maculatus: Subías 2022 .

Neotropic: Chile (Juan Fernandez Island) ( Balogh & Balogh 1987, 1988, Niedbała 1986b, 1992, 1994a, 2004a, Subías 2022, Trägårdh 1931).

Neotropical, Juan Fernandez Island, endemic maculosus Atropacarus (Atropacarus) (Niedbała, 1983)

Steganacarus (Atropacarus) maculosus Niedbała, 1983

Atropacarus maculosus: Subías 2004 .

Atropacarus (Atropacarus) maculosus: Niedbała 1986b, 1992 , 2011b, 2012b, Niedbała & Liu 2018, Subías 2022.

Atropacarus kremenitsai Shtanchaeva et Subías, 2012 syn. by Niedbała 2015a.

Atropacarus kremenitsai: Subías 2012 .

Palaearctic: Georgia, Russia (Caucasus) ( Niedbała 1983a, 1986b, 1992, 2011b, 2012b, 2015a, Subías 2022).

Western Palaearctic, Caucasus madagassica Bursoplophora Mahunka, 1994

Bursoplophora madagassica: Niedbała 2004b , Niedbała & Liu 2018, Subías 2022.

Afrotropic: Madagascar ( Mahunka 1994a, 2002, Niedbała 2004b, 2017a, 2008c, Subías 2022). Afrotropical, Madagascar, endemic madegassica Mesoplophora (Parplophora) Mahunka, 2009

Mesoplophora (Parplophora) madegassica: Liu 2016 a, Niedbała & Liu 2018 , Subías 2022.

Afrotropic: Madagascar (Mahunka 2009b, Niedbała 2017a, Subías 2022).