Anotylus schuelkei, Makranczy, 2017

Makranczy, György, 2017, Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae), Acta Zoologica Academiae Scientiarum Hungaricae 63 (2), pp. 143-262 : 252-254

Anotylus schuelkei sp. n.

( Figs 84 , 244–250 , 276 )

Typematerial – Holotype (m): CHINA: Yunnan, NujiangLisuAuton. Prefecture, Gaoligong Shan , “Cloud pass” 21 km NW Liuku, 25°58’21”N, 98°41’01”E, 3150 m, 2.IX.2009, leg. M. Schülke (CH09-22), shrubs & bamboo, littersifted (coll. SchülkeinZMHB) ; Paratypes (285): samedataasholotype (24, coll. Schülke, 2, IZAS); CHINA: Yunnan, NujiangLisu Auton. Prefecture , creekvalley 3 kmSEGongshan, 27°43’02”N, 98°41’27”E, 1450–1500 m, 5. VI.2007, leg. D.W. Wrase (20) , litter/mosssifted (3, coll. Schülke); Yunnan, NujiangLisu Auton. Prefecture , GaoligongShan, pass 21 kmNWLiuku, 25°58’22”N, 98°41’00”E, 3150 m, 9. VI.2007, leg. M. Schülke ( CH 07-26) , bamboowithshrubs, littersifted (1, coll. Schülke); Yunnan , NujiangLisuAuton. Prefecture, GaoligongShan, creekvalley 20 kmNWLiuku, 25°58’49”N, 98°41’48”E, 3000 m, 9. VI.2007, leg. D.W. Wrase (27) , bamboo, shrubs, litter sifted (11, coll. Schülke); Yunnan , NujiangLisuAuton. Prefecture, GaoligongShan, creek valley 20 km NW Liuku, 25°58’49”N, 98°41’48”E, 3000 m, 9. VI.2007, leg. M. Schülke ( CH 07- 27) , bamboo, shrubs, littersifted (13, coll. Schülke); Yunnan , NujiangLisuAuton. Prefec- ture, GaoligongShan, sidevalley 19 kmNWLiuku, 25°59’02”N, 98°42’23”E, 2730 m, 9-10. VI.2007, leg. D.W. Wrase (28) , devastatedprimaryforest, littersifted (28, coll. Schülke, 1, SMNS, 1, CNCI, 1, SDEI, 1, FMNH, 1, ISNB, 1, MNHP, 1, NIBR, 1, BMNH, 1, NKME, 1, NMPC, 1, AMNH, 1, ZMUC, 1, coll. Assing, 1, coll. Kleeberg, 3 m, 1 f, 1, HNHM); Yunnan , NujiangLisuAuton. Prefecture, GaoligongShan, sidevalley 19kmNWLiuku, 25°59’02”N, 98°42’23”E, 2730 m, 9. VI.2007, leg. M. Schülke ( CH 07-28) , devastatedprimaryforest, litter sifted (65, coll. Schülke); samebut 10. VI.2007 ( CH 07-28a) (49, coll. Schülke); Yunnan , NujiangLisuAuton. Prefecture, GaoligongShan, sidevalley 18 kmNWLiuku, 25°58’10”N, 98°42’27”E, 2590 m, 9-10. VI.2007, leg. D.W. Wrase (29) , devastatedprimaryforest, litter sifted (6, coll. Schülke); Yunnan , NujiangLisuAuton. Prefecture, GaoligongShan, “Cloud pass” 21 km NW Liuku, 25°58’21”N, 98°41’01”E, 3150 m, 2.IX.2009, leg. D.W. Wrase (22a) , shrubs, Vaccinium , bamboo, littersifted (4, coll. Schülke, 6, MHNG); Yunnan , Nujiang LisuAuton. Prefecture, GaoligongShan, “Cloudpass” 21 kmNWLiuku, 25°58’21”N, 98°41’01”E, 3150 m, 3.IX.2009, leg. M. Schülke ( CH 09-22a) , shrubsandbamboo, littersifted (10, coll. Schülke, 1, HNHM); Yunnan , NujiangLisuAuton. Prefecture, GaoligongShan, W “Cloud pass” 24 km NW Liuku, 25°59’02”N, 98°39’56.5”E, 2940 m, 3.IX.2009, leg. M. Schülke ( CH 09-24) , small cleft, wet moss & litter sifted (1, coll. Schülke); Yunnan , Nujiang Lisu Auton. Prefecture, GaoligongShanEpass 20 kmNWLiuku, 25°58’49”N, 98°41’48”E, 3000 m, 3.IX.2009, leg. D.W. Wrase (25) , creekvalleywithdevastatedprimaryforest, ferns, litter/ mosssifted (13, coll. Schülke, 2, IZAS, 7, NHMW); Yunnan , NujiangLisuAuton. Prefec- ture, Gaoligong Shan E pass 20 km NW Liuku, 25°58’49”N, 98°41’48”E, 3000 m, 3.IX.2009, leg. M. Schülke ( CH 09-25) , creekvalleywithdevastatedprimaryforest, litter & mosssifted (20, coll. Schülke).

Description – Measurements (in mm, n=10): HW = 0.60 (0.57–0.62); TW = 0.60 (0.56– 0.63); PW = 0.79 (0.73–0.81); EW = 0.79 (0.72–0.84); AW = 0.94 (0.88–1.02); HL = 0.45 (0.43– 0.47); EL = 0.10 (0.10–0.11); TL = 0.18 (0.17–0.18); PL = 0.48 (0.46–0.50); SC = 0.33 (0.31–0.35); FB = 1.33 (1.27–1.35); BL = 2.98 (2.83–3.07). Habitus as in Fig. 84 . Forebody moderately shiningbecauseofconspicuouslightcolourationandloosesculpture, abdomenveryfinely microsculpturedwithsimilarmoderatelustre. Bodysomewhatyellowishlightbrown, abdominalbasesdarker, twoblackishspotsatthebasesofsupraantennalprominencesnear circularclypealimpressionandtwosimilarimpresseddarkpitsathindvertex. Headwith anteriormarginpossessingthinrim, somewhatunevenlyarched, anteriorclypealrimvery slightlyprojectinginmiddle, supraantennalprominencesasmoderatelyelevatedoblique ridges, discimpressedneareyes. Epistomalsutureformingsubcircularimpression (inside almostunsculptured), uppersurfacewithouttactilesetae, eyessmallandslightlybulging; indorsalviewtemples (1.8× lengthofeye) alatiform, laterallyroundedandposteriorly constrictedtoformdistinctneck, however, dorsalsidenotseparatedbyoccipitalfurrow, ratherbydorsumofneckbearingmicrosculptureinsteadofpunctation. Antennomere 1 flattenedclub-shapedandsmooth, segment 2 ovoid, segment 3 club-shaped, segment 4 smallandisodiametricalal, segments 5–11 withbasaldishes, articles 6–10 slightlytransverse (1.0–1.3× broaderthanlong). Headandpronotumwithmostlyfoveatesculpture. Pronotumwithlateralportionexplanate, edgewithafewshortsetae, lateralmarginalmostevenlyarched, hindmarginbeforecornersslightlyconcave; discmediallywithapair ofshinierparallellongitudinalridges, partlyimpressedalongside. Elytrawithinsignificantmembranousposteriormarginextendingfromsuturalcornerto 3/4 ofhindmargin. Elytralepipleuralridgepresentwithup-turnedlateralalatiformportions, discdepressed; shouldersweaklydeveloped, surfacefoveolate, inbetweenpuncturesrathersmooth, apicalmarginslightlyoblique. Legsshort, pro-andmesotibiawithseveralspinuloserows, metatibiawithlongitudinalctenidiumofspinulesindistalhalf. Abdomenwithsidesmoderatelyarcuate, secondsegmentwithparatergitesstronglybroadeningposteriorly (abdomenappearingconstrictedatbase), thoseofsegmentsIII–VIIthin, mesalparatergites broad; posterioredgeoftergiteVIIwiththinpalisadefringe, medianpartofsterniteVIIas inFig. 244 , sterniteVIII ( Fig. 245 ), tergitesIX ( Fig. 246 ), X ( Fig. 247 ) andaedeagusasinFigs 248–250 , spermathecaasinFig. 276 .

Etymology – Thespeciesisnamedafterthecollectorofmostofthematerial, Michael

Schülke (Berlin), aprivatecollectorwhosecollectionnowbelongstoZMHB.

Distribution – KnownonlyfromasmallareainYunnanprovince ( China). Remarks – Thepresenceofthisspeciessuggeststhatsimilarbrachypter- ousspeciesmayexistelsewhereinthemoremountainousareasofChina.

245–247

Figures

Figs 84–85. Habitus of Anotylus species. 84 = A. schuelkei sp. n., 85 = A. bengalensis (Biswas et Sengupta, 1980)

Figs 244–250. Anotylus schuelkei sp. n., male. 244 = median part of sternite VII, 245 = sternite VIII, 246 = tergite IX, 247 = tergite X, 248 = aedeagus, frontal view, 249 = median lobe, lateral view, 250 = paramere, lateral view. Scales: 0.10 mm for Figs 244, 248–250, 0.12 mm for Figs

Figs 265–278. Spermathecae of Anotylus species: 265 = A. coonoor sp. n., 266 = A. ganapati sp. n., 267 = A. hartmanni sp. n., 268 = A. ijen sp. n., 269 = A. jambi sp. n., 270 = A. kabasi sp. n., 271 = A. linaxi sp. n., 272 = A. riedeli sp. n., 273 = A. rurukan sp. n., 274 = A. schawalleri sp. n., 275 = A. schillhammeri sp. n., 276 = A. schuelkei sp. n., 277 = A. tanator sp. n, 278 = A. topali