Anotylus schawalleri, Makranczy, 2017

Makranczy, György, 2017, Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae), Acta Zoologica Academiae Scientiarum Hungaricae 63 (2), pp. 143-262 : 248-250

Anotylus schawalleri sp. n.

( Figs 73 , 230–236 , 274 )

Typematerial – Holotype (m): MALAYSIA /BORNEO: Sabah, CrockerRangeN. P., NWKeningau, 900–1200 m [05°26‘N, 116°05‘E], 16-20.XI.1996, leg. W. Schawaller (SMNS); Paratypes (4): samedataasholotype (1 m, HNHM); Sabah, CrockerRange, 1550–1650 m, àproximitéducol (routeKotaKinabalu-Tambunan) [05°47‘55“N, 116°20‘06“E], forêt de Lithocarpus - Castanopsis , 16.V.1996, leg. D. Burckhardt & I. Löbl (27a), tamisagedebois pourri, feuillesmortesetmousses (1 m, MHNG); Sabah, CrockerRange, faceouest (W), 1600 m env., km 51-52 - route de Kota Kinabalu - Tambunan [05°50‘04“N, 116°20‘08“E], foréthumideà Lithocarpus , Castanopsis etfougèresarborescentes, 18.V.1996, leg. D. Burck- hardt & I. Löbl (30a), tamisagededébrisvegetauxdansunravinhumideetàlalisièrede laforêt (1 f, MHNG); Sabah, BanjaranCrockerMts., GunungAlabpeak, 30.IV.-27.V.1996, 1650– 1800 m [05°50‘N, 116°20‘E], leg. M. Štrba & R. Hergovits (1 m, NHMW).

Description – Measurements (in mm, n=5): HW = 0.60 (0.58–0.61); TW = 0.57 (0.56–0.59); PW = 0.82 (0.80–0.85); EW = 0.94 (0.92–0.96); AW = 0.92 (0.91–0.93); HL = 0.43 (0.42–0.44); EL = 0.14 (0.13–0.15); TL = 0.18 (0.17–0.19); PL = 0.51 (0.50–0.52); SC = 0.60 (0.59–0.61); FB = 1.61 (1.58–1.63); BL = 3.05 (2.91–3.25). HabitusasinFig. 73 . Forebodyweaklyshiningdespite sculpture, abdomenwithpuncturesofvaryingsizes, ratherdensebutshallow, indistinctly microsculptured, yetmorelustrousthanforebody. Headblackishdarkbrown, pronotum, elyraandabdomenreddishdarkbrown, occasionallylighter, mediumbrownstripealong posteriorelytralmargin, lateralpronotalmarginandanteriormarginofhead. Legs, mouthpartsandantennaereddishmediumbrown. Headwithanteriormarginpossessingrather strongrim, almosttruncateinfront, laterallystraighttosupraantennalprominences (moderatelyelevatedobliqueridges), discslightlyimpressedneareyes. Epistomalsutureformingsubcircularimpression (muchlesspunctateinside), uppersurfacewithweaktactilesetae (intratemporalis) situatedpostero-mediallyofeacheye; eyesmediumsizedandslightly bulging, indorsalviewtemples (1.3× lengthofeye) alatiform, laterallyevenlyroundedand posteriorlyconstrictedtoformdistinctneckseparatedbyweakoccipitalfurrow. Anten- nomere 1 flattenedclub-shaped, stronglysculptured, segment 2 basallybroadsubtriangular, segment 3 incrassate, segment 4 transversediscoidwithrim, segments 5–11 withbasal dishes, articles 5–10 transverse (about 1.5–1.7× broaderthanlong). Headandpronotum withmostlyalveolatesculpture. Pronotumwithlateralportionexplanate, edgealatiform withsparseshortsetae; lateralmarginunevenlyarchedwithanumberofsmall, irregularly distributeddenticles, posteriorcornersnotincised, discmediallywithtwoweakparallel longitudinalridges, pronotalsidesweaklyimpressedaroundmiddle. Elytrawithoutmembranousposteriormargin. Elytralepipleuralridgeapparentasstrongersculpturelinesin lateralalatiformportions, discwithveryslighttonoimpressionincentre; shoulderswell developed, surfaceobscurelyfoveolateandinbetweenpuncturestorulose. Legsshort, pro- andmesotibiawithseveralspinuloserows, metatibiawithlongitudinalctenidiumofspi- nulesindistalhalf. Abdomenwithsidesmoderately/ratherarcuate, secondsegmentwith paratergitesbroadeningposteriorly (abdomenappearingconstrictedatbase), thoseofseg- mentsIII–VIIthin, mesalparatergitesbroad, posterioredgeoftergiteVIIwiththinpalisade fringe, medianpartofsterniteVIIasinFig. 230 , sterniteVIII ( Fig. 231 ), tergitesIX ( Fig. 232 ), X ( Fig. 233 ) andaedeagusasinFigs 234–236 , spermathecaasinFig. 274 .

Etymology – Thespeciesisnamedafterthecollectoroftheholotype, WolfgangScha-

waller (SMNS).

Distribution – Thespeciesappearstobeendemictothehighestareasof theCrockerRangeinSabah (MalaysianBorneo).

Remarks – Thelownumberofspecimensexamineddoesnotallowagood judgementofintraspecificvariability, butexternallythespeciescanbedifficult to separate from the closely related A. borneensis .

Figures

Figs 73–75. Habitus of Anotylus species. 73 = A. schawalleri sp. n., 74 = A. borneensis (Cam- eron, 1928), 75 = A. gunung sp. n.

Figs 230–236. Anotylus schawalleri sp. n., male. 230 = median part of sternite VII, 231 = ster- nite VIII, 232 = tergite IX, 233 = tergite X, 234 = aedeagus, frontal view, 235 = median lobe, lateral view, 236 = paramere, lateral view. Scales: 0.10 mm for Figs 234–236, 0.115 mm for Fig. 233, 0.15 mm for Figs 230–232

Figs 265–278. Spermathecae of Anotylus species: 265 = A. coonoor sp. n., 266 = A. ganapati sp. n., 267 = A. hartmanni sp. n., 268 = A. ijen sp. n., 269 = A. jambi sp. n., 270 = A. kabasi sp. n., 271 = A. linaxi sp. n., 272 = A. riedeli sp. n., 273 = A. rurukan sp. n., 274 = A. schawalleri sp. n., 275 = A. schillhammeri sp. n., 276 = A. schuelkei sp. n., 277 = A. tanator sp. n, 278 = A. topali