Anotylus rurukan, , Gunung Mahawu, 1200

Makranczy, György, 2017, Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae), Acta Zoologica Academiae Scientiarum Hungaricae 63 (2), pp. 143-262 : 245-247

Anotylus rurukan sp. n.

( Figs 96 , 223–229 , 273 )

Typematerial – Holotype (m): INDONESIA / SULAWESI: N-, Tomohon , Rurukan, Gunung Mahawu [01°20‘53“N, 124°51‘46“E], 1200 m, 1-4.XII.1999, leg. A. Riedel ( SMNS) ; Paratypes (35): samedataasholotype (8 m, SMNS); N-, Tomohon , Rurukan, Gn. Mahawu, 1200 m, 30.XI.1999, leg. A. Riedel (2 m, 9 f, SMNS, 1 m, MHNG, 1 m, AMNH, 1 m, FMNH, 1 m, ZMUC, 1 m, ZMHB, 1 m, SDEI, 1 m, NMPC, 1 m, NIBR, 1 m, BMNH, 1 m, ISNB, 1 m, MNHP, 1 m, CNCI, 1 m, HNHM) ; 2 km W Tomohon, bottom of Mt. Lokon , 830–850 m, 01°21‘29“N, 124°48‘57“E, 29-31.I.2004, leg. A Skale (1 f, NHMW) ; N-, [nr.] Modoinding , Kotamobagu, Gn. Ambang [00°45‘27“N, 124°25‘23“E], 1450 m, 6.XII.1999, leg. A. Riedel

(1 m, SMNS); Umg. Modoinding, NEKotamobagu, 19.IV.1992, leg. M. Jäch (12), various streams, insideforestandataforestpool (1, NHMW) .

Description – Measurements (in mm, n=10): HW = 0.60 (0.53–0.64); TW = 0.57 (0.51– 0.61); PW = 0.86 (0.75–0.92); EW = 0.90 (0.84–0.97); AW = 0.96 (0.88–1.03); HL = 0.47 (0.43– 0.50); EL = 0.12 (0.11–0.13); TL = 0.18 (0.16–0.19); PL = 0.48 (0.41–0.51); SC = 0.54 (0.49–0.57); FB = 1.54 (1.42–1.62); BL = 3.02 (2.77–3.20). Habitus as in Fig. 96 . Forebody weakly shining becauseofsculpture; abdomenwithdensepuncation, large (butshallow), apicallyfiner punctures, fromtergiteVIonwardsabdomenveryfinelycoriaceousmicrosculptured, morelustrousthanforebody. Forebody, legsandabdomenreddishdarkbrown, except explanatepartofpronotumandclypealarea (bothfortransparency), these, mouthparts andantennaereddishmediumbrown. Headrathertransverse (clypealarealessprojecting forward), withathinrimonanteriormargin, truncateinmiddle, veryslightlyangledalso atsupraantennalprominences, theseasmoderatelyelevatedobliqueridges, discslightly impressedneareyes. Epistomalsutureformingsubcircularimpression (lesspunctured inside), uppersurfacewithouttactilesetae, eyesmoderatelysmall, veryslightlybulging fromsidelineofhead; indorsalviewtemples (1.5× lengthofeye) alatiform, laterallyimperfectlyrounded (withoutbeingangled) andposteriorlyconstrictedtoformdistinctneck, notseparateddorsallybyoccipitalfurrow, onlybyneckbeingmicrosculpturedratherthan punctured. Antennomere 1 club-shaped, finelysculptured, segment 2 elongateoval, seg- ment 3 club-shaped, segment 4 smallandveryslightlytransverse, segments 5–11 with basaldishes, articles 6–10 transverse (about 1.3× broaderthanlong), segments 9–10 bulk- ierthanpreceding. Headandpronotumwithmostlylacunosesculpture. Pronotumwith lateralportionexplanateandslightlyup-turned, edgealatiformwithsparseshortsetae; lateralmarginunevenlyarchedwithanumberofsmall, irregularlydistributeddenticles, posteriormarginbroadlyincisedbeforecorners, discmediallywithtwoparallellongitudinalridges, impressedalongsidethem, laterallyimpressedaroundmiddleofsides. Elytra withmembranousposteriormarginextendingfromsuturalcornerto 3/4 ofhindmargin. Elytralepipleuralridgepresentwithup-turnedlateralalatiformportions, disclongitudinallyimpressedalongside; shoulderswelldeveloped, surfacefoveolate, inbetweenpunc- turestorulose. Legsshort, pro-andmesotibiawithseveralspinuloserows, metatibiawith longitudinalctenidiumofspinulesindistalhalf. Abdomenwithsidesratherarcuate, sec- ondsegmentwithparatergitesstronglybroadeningposteriorly (abdomenappearingcon- strictedatbase), thoseofsegmentsIII–VIIthin, mesalparatergitesbroad, posterioredgeof tergiteVIIwiththinpalisadefringe, medianpartofsterniteVIIasinFig. 223 , sterniteVIII ( Fig. 224 ), tergitesIX ( Fig. 225 ), X ( Fig. 226 ) andaedeagusasinFigs 227–229 , spermatheca asinFig. 273 .

Etymology – Thespecificepithetisfromthetypelocality, Rurukan (nouninapposition).

Distribution – SofaritisonlyknownfromnorthernSulawesi ( Indonesia). Remarks – Thisspeciesiscloselyrelatedto A. tanator (alsofromSulawesi, butfromthesouthernpart).

225, 0.14 mm for Fig. 224

Figures

Figs 96–97. Habitus of Anotylus species. 96 = A. rurukan sp. n., 97 = A. tanator sp. n.

Figs 223–229. Anotylus rurukan sp. n., male. 223 = median part of sternite VII, 224 = sternite VIII, 225 = tergite IX, 226 = tergite X, 227 = aedeagus, frontal view, 228 = median lobe, lateral view, 229 = paramere, lateral view. Scales: 0.10 mm for Figs 226–229, 0.12 mm for Figs 223,

Figs 265–278. Spermathecae of Anotylus species: 265 = A. coonoor sp. n., 266 = A. ganapati sp. n., 267 = A. hartmanni sp. n., 268 = A. ijen sp. n., 269 = A. jambi sp. n., 270 = A. kabasi sp. n., 271 = A. linaxi sp. n., 272 = A. riedeli sp. n., 273 = A. rurukan sp. n., 274 = A. schawalleri sp. n., 275 = A. schillhammeri sp. n., 276 = A. schuelkei sp. n., 277 = A. tanator sp. n, 278 = A. topali