Anotylus linaxi, Makranczy, 2017

Makranczy, György, 2017, Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae), Acta Zoologica Academiae Scientiarum Hungaricae 63 (2), pp. 143-262 : 218-220

Anotylus linaxi sp. n.

( Figs 77 , 167–173 , 271 )

Type material – Holotype (m): CHINA: N-Yunnan, Lijiang Naxi Aut. Co., 3 km NW Yongsheng, 53 km WSW Lijiang, 1950–2000 m, 26°41.8’N, 100°43.1’E, 14.VIII.2003, leg. M. Schülke (C03-03), SEslope, secondarybroadleavedforest (coll. SchülkeinZMHB); Para- types (25): same data as holotype (1 m, 8, coll. Schülke, 1, AMNH, 1, MNHP, 1, ISNB, 1, NIBR, 1 m, NHMW, 1 m, SDEI, 1 f, BMNH, 1 m, FMNH, 1 m, HNHM), same but 1990 m, leg. A. Smetana (C122), siftingoflayersofleaflitterandotherdebrisundertallbushesopen area (3, coll. Smetana / NSMT, 1 CNCI, 1 m, ZMUC, 1 m, NMPC); THAILAND: Chiang Mai, DoiInthanon, ravintrèshumide, 1650 m, 7.XI.1985, leg. D. Burckhardt & I. Löbl (16), tamisagededébrisvégétauxetdechampignonsprèsd‘unruisselet (1 f, MHNG).

Description – Measurements (in mm, n=10): HW = 0.59 (0.57–0.62); TW = 0.58 (0.55– 0.62); PW = 0.77 (0.73–0.80); EW = 0.86 (0.81–0.90); AW = 0.87 (0.82–0.93); HL = 0.47 (0.44– 0.50); EL = 0.13 (0.13–0.15); TL = 0.19 (0.19–0.20); PL = 0.50 (0.49–0.54); SC = 0.58 (0.56–0.61); FB = 1.65 (1.61–1.77); BL = 3.23 (2.85–3.55). Habitus as in Fig. 77 . Forebody weakly shining 0.11 mm for Figs 167, 169 , 0.12 mm for Fig. 168 becauseofsculpture, abdomenwithlargerbutindistinctlyborderedandshallowpunc- tures, tracesofmicrosculpture, somewhatmorelustrous. Bodyreddishmediumbrown, head (exceptclypealarea) oftendarker, darkbrown, middleoftergitebasesoftenblackish aswellasmostofantennomere 1. Headwithanteriormarginpossessingrim (weakerin middle), evenlyarchedbutslightlyangledatsupraantennalprominences, theseasmod- eratelyelevatedobliqueridges, rimcontinuingposteriadalonginnerborderofeyes, disc slightlyimpressedbesideit. Epistomalsutureformingcircularimpression (lesssculptured inside), uppersurfacewithweaktactilesetae (intratemporalis) situatedpostero-medially ofeacheye. Eyesmedium-sizedandslightlybulging, indorsalviewtemples (1.4× length ofeye) alatiform, laterallyroundedandposteriorlyconstrictedtoformdistinctneck, separatedbytransversal, weakoccipitalfurrow. Antennomere 1 flattenedclub-shaped andstronglymicrosculptured, segment 2 ovalbutbroadatbase, segment 3 club-shaped, segment 4 smallandslightlytransverse, segments 5–11 withbasaldishes, articles 6–10 transverse (about 1.5× broaderthanlong), apexofsegment 11 lighter. Headwithareolate- rugose, pronotumwithlacunosesculpture. Pronotumwithlateralportionexplanateand slightlyup-turned, edgealatiformwithverysparseshortsetae; lateralborderconsistingof twolongerstraightsectionsatanangleconnectedbyshortconcavesectionwiththinmar- ginalridgeandanumberofsmall, indistinctdenticles, posteriormarginevenlyarched; discmediallywithtwonearlyparallelandposteriorlyconfluentlongitudinalridges, posteriorlyimpressedalongsidethem, laterallysomewhatimpressedtowardssidemargin. Elytrawiththinmembranousposteriormarginextendingfromsuturalcornerto 3/4 of hindmargin. Elytralepipleuralridgeapparentasstrongersculpturelinesinlateralalatiformportions, discwithveryslighttonoimpressionincentre; shoulderswelldeveloped, surfaceobscurelyfoveolateandinbetweenpuncturestorulose. Legsshort, pro-andmesotibiawithseveralspinuloserows, metatibiawithlongitudinalctenidiumofspinulesin distalhalf. Abdomenwithsidesmoderatelyarcuate, secondsegmentwithparatergites broadeningposteriorly (abdomenslightlyconstrictedatbase), thoseofsegmentsIII–VII thin, mesalparatergitesbroad; posterioredgeoftergiteVIIwiththinpalisadefringe, medianpartofsterniteVIIasinFig. 167 , sterniteVIII ( Fig. 168 ), tergitesIX ( Fig. 169 ), X ( Fig. 170 ) andaedeagusasinFigs 171–173 , spermathecaasinFig. 271 .

Etymology – The specific epithet is a combination of the first letters of ‘Lijiang’ and

‘Naxi’, fromthelocalitylabel (nouninapposition).

Distribution – The know distribution is limited to Yunnan ( China) and ChiangMai ( Thailand).

Remarks – The species is remarkably similar to A. reitteri and A. chinensis , differsonlyinitssmallersize, lesssculpturedbodysurfaceandaslightdiffer- enceinpronotalshape (lessroundedsideswithacharacteristic “step” before the middle).

Figures

Figs 76–77. Habitus of Anotylus species. 76 = A. coonoor sp. n., 77 = A. linaxi sp. n.

Figs 167–173. Anotylus linaxi sp. n., male. 167 = median part of sternite VII, 168 = sternite VIII, 169 = tergite IX, 170 = tergite X, 171 = aedeagus, frontal view, 172 = median lobe, lateral view, 173 = paramere, lateral view. Scales: 0.09 mm for Figs 170, 0.10 mm for Figs 171–173,

Figs 265–278. Spermathecae of Anotylus species: 265 = A. coonoor sp. n., 266 = A. ganapati sp. n., 267 = A. hartmanni sp. n., 268 = A. ijen sp. n., 269 = A. jambi sp. n., 270 = A. kabasi sp. n., 271 = A. linaxi sp. n., 272 = A. riedeli sp. n., 273 = A. rurukan sp. n., 274 = A. schawalleri sp. n., 275 = A. schillhammeri sp. n., 276 = A. schuelkei sp. n., 277 = A. tanator sp. n, 278 = A. topali