Anotylus hartmanni, Makranczy, 2017

Makranczy, György, 2017, Review Of The Anotylus Cimicoides Species Group (Coleoptera: Staphylinidae: Oxytelinae), Acta Zoologica Academiae Scientiarum Hungaricae 63 (2), pp. 143-262 : 207-209

Anotylus hartmanni sp. n.

( Figs 82 , 132–138 , 267 )

Typematerial – Holotype (m): NEPAL: ManasluMts., SE-slope, WGupchiDanda, 2200–2300 m [28°08‘37“N, 84°44‘42“E], 28.V.2006, leg. J. Schmidt, sifting leaf litter in for- estremainsalongmountaincrest ( NKME) ; Paratypes (41): samedataasholotype (1 m, NHMW) ; ManasluMts., DudhPokhariLekh, belowHelamPokhari, 2000 m [28°11‘N, 84°34‘E], 22.IV.2003, leg. J. Schmidt, sifting forest litter (2 m, 3 f, 22, NKME, 1, MNHP, 1, ZMUC, 1, ISNB, 1, NMPC, 1, CNCI, 1, FMNH, 1, AMNH, 1, MHNG, 1, SDEI, 1, SMNS, 1, NIBR, 1 m, BMNH, 1 m, HNHM) ; NEPAL: Manaslu Mts. , Dudh Pokhari Lekh , upper Phulinagiri, Madi, 2500 m [abovePhulingiriMadi, 28°13‘N, 84°34‘E], 19-21.IV.2003, leg. J. Schmidt, siftingforestlitter (1, NKME) .

Description – Measurements (in mm, n=10): HW = 0.63 (0.60–0.65); TW = 0.65 (0.61– 0.68); PW = 0.90 (0.85–0.97); EW = 0.90 (0.84–0.95); AW = 0.97 (0.90–1.03); HL = 0.48 (0.45– 0.51); EL = 0.11 (0.10–0.12); TL = 0.21 (0.20–0.23); PL = 0.57 (0.53–0.60); SC = 0.48 (0.45–0.50); FB = 1.58 (1.49–1.65); BL = 3.30 (2.61–3.50). Habitus as in Fig. 82 . Forebody moderately shiningdespitesculpture, abdomenwithfaintcoriaceousmicrosculpture, shallowlypunc- tured, aslustrousasforebody. Bodyreddishdarkbrownexceptabdomen, anteriormargin ofhead, lateralmarginofpronotum, antennaeandmouthpartsmediumtodarkbrown. Headwithanteriormargintruncateinmiddle, obliquetosupraantennaltubercles (asmod- eratelyelevatedobliqueridges) wheregentlyangled (broadlyrounded) again, possessing slightrim, discslightlyimpressedneareyes. Epistomalsutureformingsubcircularimpression (lesspuncturedinside), uppersurfacewithouttactilesetae, eyessmallandrather flat; indorsalviewtemples (1.9× lengthofeye) alatiform, laterallybulgingandrounded, posteriorlyconstrictedtoformdistinctneck, however, dorsalsidenotseparatedbyoccipi- talfurrow, ratherbydorsumofneckbearingmicrosculptureinsteadofpunctation. Anten- nomere 1 flattenedclub-shapedandsmooth, segment 2 basallybroadovoid, segment 3 club-shaped, segment 4 smallandisodiametrical, segments 5–11 withbasaldishes, articles 6–10 slightlytransverse (about 1.4× broaderthanlong). Headandpronotumwithmostly lacunosesculpture. Pronotumwithlateralportionexplanateandslightlyup-turned, edge withafewshortsetae, lateralborderratherevenlyarched, withanumberofsmall, irregularlydistributedtubercles, posteriormarginincisedbeforecorners; discmediallyrather convex, withtwoposteriorlyconfluentlongitudinalridges, impressedalongsidethem, laterallyimpressednearsidemargins. Elytrawithmembranousposteriormarginextendingfromsuturalcornertolateral 5/6 ofhindmargin, asmallincision- likethinningbefore outercorner. Elytralepipleuralridgepresentwithup-turnedlateralalatiformportions, discdepressed; shouldersmoderatelydeveloped, surfacefoveolate, inbetweenpunctures torulose. Legsshort, pro-andmesotibiawithseveralspinuloserows, metatibiawithlon- gitudinalctenidiumofspinulesindistalhalf. Abdomenwithsidesweaklyarcuate, second segmentwithparatergitesstronglybroadeningposteriorly (abdomenappearingconstrict- edatbase), thoseofsegmentsIII–VIIthin, mesalparatergitesbroad. Posterioredgeof tergiteVIIwiththinpalisadefringe, medianpartofsterniteVIIasinFig. 132 , sterniteVIII ( Fig. 133 ), tergitesIX ( Fig. 134 ), X ( Fig. 135 ) andaedeagusasinFigs 136–138 , spermatheca asinFig. 267 .

Etymology – ThespeciesisnamedafterMatthiasHartmann (NKME), whocontributedsignificantlytothebetterknowledgeofthefaunaoftheHimalayas, particularlyNepal.

Distribution – OnlyknownfromtheManaslumountainrangeinWestern Nepal.

Remarks – Habitatnotesfromthecollector (JoachimSchmidt) forthe 40 paratypescollectedin 2003: Forestremnantsneartemporarystreams; species richforestsonbothcollectingsites, mainlyevergreentreesandshrubs, but markedlyinfluencedbytreecuttingandcow/yak/sheep/goatgrazing.

Figures

Figs 82–83. Habitus of Anotylus species. 82 = A. hartmanni sp. n., 83 = A. schillhammeri sp. n.

Figs 132–138. Anotylus hartmanni sp. n., male. 132 = median part of sternite VII, 133 = ster- nite VIII, 134 = tergite IX, 135 = tergite X, 136 = aedeagus, frontal view, 137 = median lobe, lateral view, 138 = paramere, lateral view. Scales: 0.10 mm for Figs 136–138, 0.125 mm for Figs 134–135, 0.14 mm for Figs 132–133

Figs 265–278. Spermathecae of Anotylus species: 265 = A. coonoor sp. n., 266 = A. ganapati sp. n., 267 = A. hartmanni sp. n., 268 = A. ijen sp. n., 269 = A. jambi sp. n., 270 = A. kabasi sp. n., 271 = A. linaxi sp. n., 272 = A. riedeli sp. n., 273 = A. rurukan sp. n., 274 = A. schawalleri sp. n., 275 = A. schillhammeri sp. n., 276 = A. schuelkei sp. n., 277 = A. tanator sp. n, 278 = A. topali