Landouria sewuensis, Nurinsiyah & Neiber & Hausdorf, 2019
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publication ID |
https://doi.org/ 10.5852/ejt.2019.526 |
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publication LSID |
lsid:zoobank.org:pub:3FFC527D-E9CA-4D9F-BF59-076FA60171AC |
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DOI |
https://doi.org/10.5281/zenodo.11238077 |
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persistent identifier |
https://treatment.plazi.org/id/2E324442-51A9-4897-9635-22666EF0E037 |
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taxon LSID |
lsid:zoobank.org:act:2E324442-51A9-4897-9635-22666EF0E037 |
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treatment provided by |
Plazi |
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scientific name |
Landouria sewuensis |
| status |
sp. nov. |
Landouria sewuensis View in CoL sp. nov.
urn:lsid:zoobank.org:act:2E324442-51A9-4897-9635-22666EF0E037
Figs 23 View Figs 15–24 , 37 View Figs 29–47 , 53 View Figs 52–55 , 58 View Figs 56–59 ; Tables 3–4 View Table 3 View Table 4
Landouria cf. rotatoria – Köhler et al. in press View in CoL : 6 (in part). (not Pfeiffer, 1842)
Diagnosis
Landouria sewuensis sp. nov. is characterized by a brownish-corneous, sharply keeled shell with a wide umbilicus, a penis that is divided into a long, broad distal and a short, narrow proximal part with a globular bulge at its proximal end, a broad flagellum with two rows of broad tubercles and a globular bulge in the distal part of the vagina.
Etymology
Landouria sewuensis sp. nov. is named after its occurrence in the Gunung Sewu karst region.
Material examined
Holotype
INDONESIA • holotype (det. anat.); Gunung Kidul , Tepus, Mbajo Hill, natural forest; 8°08′33″ S, 110°40′59″ E; 249 m a.s.l.; 8 Aug. 2014; A.S. Nurinsiyah, F.L.H. Irsyad, F.J. Sari and E. Nurlaela leg.; D = 15.1 mm, H = 7.1 mm; MZB 19255 .
GoogleMapsParatypes
INDONESIA – Central Java • many spec.; Yogyakarta, Gunung Kidul, Kemadang: Pantai Baron ; 8°07′43″ S, 110°32′54″ E; 20 m a.s.l.; ZMH 140024 View Materials • 2 spec. (det. anat.); Gunung Kidul, Tepus, Cemara Udang Hill , secondary forest; 8°08′55″ S, 110°36′37″ E; 39 m a.s.l.; MZB 19261 GoogleMaps • 1 spec.; same data as for preceding; ZMH 133418 View Materials GoogleMaps • many spec.; Gunung Kidul, Tepus, rock above Poktunggal Beach ; 8°09′23″ S, 110°37′22″ E; 10 m a.s.l.; ZMH 140025 View Materials GoogleMaps • 1 spec. (det. anat.); Gunung Kidul, Tepus, Pego Hill , agroforestry; 8°06′10″ S, 110°37′50″ E; 230 m a.s.l.; MZB 19259 GoogleMaps • 1 spec. (det. anat.); Gunung Kidul, Tepus, Pego Hill , agroforestry; 8°06′10″ S, 110°37′51″ E; 244 m a.s.l.; MZB 19258 GoogleMaps • 1 spec.; Gunung Kidul, Tepus, near Poktunggal Beach , agroforestry; 8°08′42″ S, 110°37′52″ E; 137 m a.s.l.; MZB 21269 GoogleMaps • 3 spec. (det. anat.); same data as for preceding; MZB 19260 GoogleMaps • 3 spec.; same data as for preceding; ZMH 79891 GoogleMaps • 1 spec. (det. anat.); Gunung Kidul, Tepus, Mbajo Hill , natural forest; 8°08′32″ S, 110°40′58″ E; 281 m a.s.l.; MZB 19257 GoogleMaps • 1 spec.; same data as for preceding; ZMH 133419 View Materials GoogleMaps • 1 spec. (det. anat.); Gunung Kidul, Tepus, Mbajo Hill , natural forest; 8°08′33″ S, 110°40′59″ E; 258 m a.s.l.; MZB 19256 GoogleMaps .
GoogleMapsDescription
SHELL (Fig. 58; Tables 3–4 View Table 3 View Table 4 ). Depressed conical, with 5.25–6 hardly convex whorls; protoconch almost smooth; teleoconch with fine, irregular wrinkles; apical side with indistinct incised spiral lines, umbilical side with more distinct spiral lines; scaly processes, if present, at periphery; tuberculate around umbilicus; brownish-corneous; body whorl sharply keeled; aperture rounded rhombic; upper insertion of peristome slightly descending; peristome expanded, reflexed and slightly thickened; umbilicus slightly eccentric, wide, comprising 30–40% of shell diameter, hardly obscured by columellar edge.
GENITALIA ( Figs 23 View Figs 15–24 , 37 View Figs 29–47 ; Table 4 View Table 4 ). Atrium short; penis long, distal part broad, shorter proximal part narrow, with a globular swelling at proximal end; without penis coecum; membranaceous sheath around distal part of penis connected with proximal part of epiphallus by tissue; penial retractor runs from diaphragm to middle of cylindrical epiphallus; flagellum moderately long, broad, with two rows of broad tubercles at margin, tapering towards its proximal end; without node; vas deferens narrow and long; vagina long, distally globularly dilated, constriction towards its middle, becoming broader again towards its proximal end; oviduct short, broad, proximally tapering; peduncle of bursa copulatrix subdivided into a broader, cylindrical distal part and a narrower, longer proximal part, ending in an elongate oval bursa beside albumen gland. Right ommatophoral retractor runs between penis and vagina.
Remarks
The shell of L. sewuensis sp. nov. cannot be reliably distinguished from those of L. sukoliloensis sp. nov. (Fig. 59) or L. intumescens ( Figs 72–74 View Figs 72–75 ). Landouria sewuensis sp. nov. differs from these species in the division of the penis into a long, broad distal and a short, narrow proximal part with a globular bulge at its proximal end, the broad flagellum with two rows of broad tubercles and the globular bulge in the distal part of the vagina.
Landouria sewuensis sp. nov. occurs sympatrically with L. tholiformis sp. nov. (Fig. 51). For differences, see remarks under that species.
A sequence of L. sewuensis sp. nov. was included in the phylogenetic analysis of Köhler et al. (in press: fig. 2; ZMH 79891; erroneously labelled “ Sumatra ”) under the name L. cf. rotatoria (not Pfeiffer, 1842).
Distribution
Landouria sewuensis sp. nov. is endemic to the Gunung Sewu karst region in Gunung Kidul Regency (Fig. 53).
No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.
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Landouria sewuensis
| Nurinsiyah, Ayu Savitri, Neiber, Marco T. & Hausdorf, Bernhard 2019 |
Landouria cf. rotatoria – Köhler et al. in press
| Köhler et al. in press |