EUGLANDININAE Baker, 1941

Albesa, Joaquín & Neubauer, Thomas A., 2023, Shifting shell morphology in a Late Miocene-Pliocene land snail species lineage (Gastropoda: Stylommatophora: Spiraxidae), with the description of a new species, Geodiversitas 45 (21), pp. 643-657 : 648

Subfamily EUGLANDININAE Baker, 1941

REMARKS

The genera Palaeoglandina , Pseudoleacina and Trachyglandina are traditionally included in Oleacinidae ( Wenz 1923; Nordsieck 2014; Harzhauser & Neubauer 2021). However, this systematic placement is based on an outdated concept that unites true oleacinids and European species presently considered to belong in Spiraxidae (compare e.g. Schileyko 2000 vs Hausdorf & Solvery 2021). Nordsieck (2014) considered the genera Palaeoglandina and Pseudoleacina to belong in Euglandininae , which is today ranked as a subfamily of Spiraxidae (see also Salvador 2013). We follow that concept, which is congruent with the similarity of the fossil species and modern representatives of the subfamily, such as Euglandina and Poiretia ( Schileyko 2000) . The monotypic genus Trachyglandina is here only tentatively referred to the Euglandininae , since it is based solely on a fragment; until further material of the type species, Trachyglandina dietrichii Pfeffer, 1930 , is discovered both genus and type species should be considered nomina dubia.

The revised placement of the three genera is also supported by the geographic distribution of modern Euglandininae , some of which occur in the Mediterranean region and the Caucasus (e.g. Schileyko 2000; Welter-Schultes 2012; Helwerda 2015), while true Oleacinidae are restricted to the Americas ( Schileyko 2000; Barker & Efford 2004) (note that some of these authors included the relevant genera in Oleacinidae ).