Coenoptychus Simon, 1885

Paul, Jimmy, Sankaran, Pradeep M., Sebastian, Pothalil A. & Joseph, Mathew M., 2018, A review of the wasp mimicking spider genus Coenoptychus Simon, 1885 (Araneae: Corinnidae: Castianeirinae), Zootaxa 4413 (1), pp. 163-172 : 164-165

publication ID	https://doi.org/ 10.11646/zootaxa.4413.1.6
publication LSID	lsid:zoobank.org:pub:EED9AB62-B396-41F6-B294-AE965BA055B0
DOI	https://doi.org/10.5281/zenodo.5979928
persistent identifier	https://treatment.plazi.org/id/03B087C4-FF8B-5516-96A4-40C0FED8FF5B
treatment provided by	Plazi
scientific name	Coenoptychus Simon, 1885
status

Treatment

Coenoptychus Simon, 1885 View in CoL

Coenoptychus Simon, 1885: 36 View in CoL ; Simon 1897: 174; Majumder & Tikader, 1991: 145; Coenoptychus Deeleman-Reinhold, 2001: 329 View in CoL .

Onychocryptus Karsch, 1892: 295 .

Diagnosis. The genus Coenoptychus seems most similar to Graptartia in their shared resemblance of velvet ants, but can be distinguished by the following combination of features: rectangular eye arrangement and recurved PER ( Fig. 4A View FIGURE4 , Haddad 2004: figs 2‒3) vs circular eye arrangement and procurved PER in Graptartia ( Haddad 2004: figs 1, 19); carapace with less narrowed ocular region ( Figs 2A, C View FIGURE 2 , 4E View FIGURE4 ) vs strongly narrowed ocular region in Graptartia ( Fig. 4D View FIGURE4 ); dorsal abdominal setae feathery ( Haddad 2004: figs 11‒12) vs dorsal abdominal setae clavate in Graptartia ( Haddad 2004: fig. 10); three teeth on cheliceral promargin vs two in Graptartia ; lack of RTA ( Fig. 3A‒C View FIGURE 3 ; Haddad 2004: figs 27‒28, 32‒33) vs a small hook-shaped RTA in Graptartia (cf. Haddad 2004: figs 22‒ 23); distally pointed apical part of cymbium ( Fig. 3A‒C View FIGURE 3 ; Haddad 2004: figs 27‒28, 32‒33) vs prolaterally pointed and retrolaterally blunt apical part of cymbium in Graptartia ( Haddad 2004: figs 22‒23); corkscrew-shaped embolus directed straight or prolaterally ( Fig. 3C View FIGURE 3 ; Haddad 2004: figs 13‒14, 27‒28, 32‒33) vs clamp-shaped embolus in Graptartia ( Haddad 2004: figs 22‒23); reniform spermathecae I ( Fig. 3E‒F View FIGURE 3 ; Haddad 2004: figs 25‒26, 30‒31) vs narrow in Graptartia ( Haddad 2004: figs 20‒21).

Type species: Coenoptychus pulcher Simon, 1885 , by monotypy.

Description. Small to medium spiders (males 3.8‒6.0, females 3.95‒12.0) in length. Carapace dark reddish to orange, with dark grey borders, papillate, scattered with white long and feathery setae, medially broad, slightly elevated from eye region towards rear, highest at two-thirds its length, fovea distinct or indistinct. Eye field dark, MOQ trapezoid, AER procurved, PER recurved ( Fig. 4A View FIGURE4 , Haddad 2004: figs 2‒3). Chelicerae orange to grey, with black setae dorsally, promargin with three teeth, retromargin with two teeth, fangs baso-promarginally with modified long black setae in C. pulcher females and possibly found in C. mutillicus comb. nov. and C. tropicalis comb. nov. females ( Fig. 4G‒H View FIGURE4 ). Endites straight to slightly depressed laterally, longer than wide. Labium wider than long, with few black setae. Sternum reddish to orange, papillate, covered with feathery setae intermingled with long and short setae. Leg formula 4123, legs spinose, scattered feathery setae on legs, scopulae weak, comprising two rows of setae, leg I femur uniformly dark coloured, femora II‒IV with dark grey annulations. Abdomen oval, covered with short, black feathery setae, scattered long white and grey setae and short white feathery setae that form the pattern, dorsal scutum entire, venter covered with short and long black setae interspersed with black feathery setae. Spinnerets closely grouped together, ALS largest and subconical. Male palp segments pale orange to red-brown, without retrolateral tibial apophysis, tibia prolaterally with one pair of spines ( Fig. 3A View FIGURE 3 ; Haddad 2004: figs 27, 32); cymbium broad, gradually narrowing apically, with dense mat of tiny setae on dorsum ( Figs 3A, C View FIGURE 3 ); tegulum pear-shaped, with stump-like apical embolus base; subtegulum small, partly visible retrolaterally; embolus spiral with 1.5‒2.5 turns ( Figs 3A‒D View FIGURE 3 ; Haddad 2004: figs 13‒14, 27‒28, 32‒33). Female epigyne domed plate-like, reddish-brown, copulatory openings in circular or semi-circular ridges ( Figs 2G View FIGURE 2 , 3E View FIGURE 3 ; Haddad 2004: figs 25‒26, 30‒31), spermathecae I kidney-shaped, spermathecae II round to subtriangular ( Figs 2H View FIGURE 2 , 3F View FIGURE 3 ; Haddad 2004: figs 25‒26, 30‒31). All species mimic velvet ants ( Figs 1A‒F View FIGURE 1 , 2A, C View FIGURE 2 , 4E View FIGURE4 ).