Rubus gremlii, Focke

Tutin, T. G., Heywood, V. H., Burges, N. A., Moore, D. M., Valentine, D. H., Walters, S. M. & Webb, D. A., 1981, Flora Europaea. Volume 2. Rosaceae to Umbelliferae, Cambridge University Press : 18-19

49. R. gremlii Focke , Syn. Rub. Germ. 266 (1877)

(R. colemannii subsp. gremlii (Focke) Sudre ).

Stems procumbent or climbing, more or less hairy, with straight, yellow prickles. Leaves pedate; leaflets 3-5, glabrescent on both surfaces; terminal leaflet longstalked, ovate or ovate-oblong, narrowed to a long point, rounded or cordate at base, unevenly serrate. Inflorescence rather long and narrow, often leafy; axis and pedicels densely pubescent with patent hairs and with small acicular prickles; glands various. Sepals deflexed, tomentose; petals narrowly obovate-cuneate, pubescent, pinkish or white; stamens often much exceeding the styles. C. Europe; Britain. Au Br Cz Ge He Hu Ju Rm. Sudre breaks up the complex species R. colemannii Bloxam in Kirby, FI. Leicest. ed. 2, 38 (1850) into several subspecies. Other species, listed below, link this complex with species in Ser. Radulae :

R. balatonicus Borbâs, Result. Wiss. Erforsch. Balaton 2(2): 146 (1907). Hu.

R. beckii Halâcsy , Verh. Zool.- Bot. Ges. Wien 35: 663 (1886). Au Cz.

R. chloroclados Sabr., Österr. Bot. Zeitschr. 41: 413 (1891). Cz Rm.

R. clusii Borbâs ex Sabr. , Erdész. Lapok 1885: 104 (1885). Au Cz Ju Rm. Sudre and Nyârâdy make this species synonymous with 49.

R. condensatus P. J. Mueller , Flora (Regensb.) 41: 167 (1858). Au Br Ga Ge He Hu.

R. diminutus Gâyer in Jâv., Magyar FI. 495 (1924). Hu.

R. eriostachys P. J. Mueller & Lefèvre, Pollichia 16-Yl: 225 (1859). Au Be Br Ga Hu.

R. ferox Vest ex Tratt. , Rosac. Monogr. 3: 40 (1823) ( R. apum Fritsch , R. lasiaxon Borbâs ex Waisb. ). Au Ge Hu.

R. grandiflori^ E. I. Nyârâdy in Sävul., FI. Rep. Pop. Romàne 4: 916 (1956). Rm.

R. gratiosus P. J. Mueller & Lefèvre, Pollichia 16-17: 153 (1859). Be Cz Ga Hu.

R. gremblichii Halâcsy , Österr. Bot. Zeitschr. 40: 433 (1890). Au.

R. halacsyi Borbâs ex Halâcsy , Verh. Zool.-Bot. Ges. Wien 35: 666 (1886). Au Cz Hu.

R. hebecaulis Sudre, Bull. Assoc. Fr. Bot. 3: 101 (1900) ( R. hebeticaulis auct.). Au Br Cz Ga Ge He Ho Hu Po Rm.

R. helveticus Gremii , Beitr. Fl. Schweiz 36 (1870). Au Br Ge He.

R. indotatus Gremii , Österr. Bot. Zeitschr. 21: 128 (1871). Au He.

R. inopacatus P. J. Mueller & Lefèvre, Pollichia 16-17: 117 (1859). Au Br Cz G a Hu.

R. joannis G. Beck , Fl. Nieder-Österr. 2(1): 736 (1892). Au.

R. laetecoloratus E. I. Nyârâdy in Sävul., Fl. Rep. Pop. Romàne 4: 914 (1956). Rm.

R. morifolius P. J. Mueller , Flora (Regensb.) 41: 164 (1858). Au Cz Ge Hu.

R. mulleri Lefèvre , Pollichia 16-17: 180 (1859). Au Be Br Ga He Ho Hu Rm.

R. podophylloides Sudre, Bull. Soc. Étud. Sci. Angers 31: 105 (1902). A u C z G a H e H u.

R. porphyrantherus Hormuzaki, Mem. Sec/. $ti. (Acad. Romand) ser. 3, 2: 290 (1925). Au.

R. rariglandulosus E. I. Nyârâdy in Sävul., Fl. Rep. Pop. Romane 4: 916 (1956). Rm.

R. salisburgensis Focke , Syn. Rub. Germ. 280 (1877). Au Br Cz Ga Ge He Hu.

R. schmidelyanus Sudre, Bull. Soc. Bot. Fr. 51: 21 (1904). 2n = 35. Au Br Cz Ga Ge He Hu Po.

R. serratulifolius Sudre, Compt. Rend. Congr. Soc. Sav. (Sci.) 1908: 206 (1909). Au Be Br Ga Ge.

R. subcoriaceus E. I. Nyârâdy in Sävul., Fl. Rep. Pop. Romàne 4: 916(1956). Rm.

R. taeniarum Lindeb. , Nov. FI. Scand. 5 (1858) ( R. infestus auct., non Weihe ex Boenn.). Be Br Cz Da Ge?Ho Su.

R. teretiusculus Kaltenb. , FI. Aachen. Beck. 2: 282 (1844). Au Be Br Ge?Ho Hu.

Series Radulae Focke. Stems rough and tuberculate; prickles scattered and very unequal, quite distinct from the smaller pricklets. Inflorescence-axis with glandular and eglandular hairs, the glandular hairs mostly not longer than the eglandular.