Eniclases Waterhouse, 1879

Bocek, Matej & Adamkova, Kristyna, 2019, New species of Moluccan trichaline net-winged beetles, with remarks on the phylogenetic position and distribution of Schizotrichalus (Coleoptera: Lycidae: Metriorrhynchinae), Zootaxa 4623 (2), pp. 341-350 : 344-345

Eniclases Waterhouse, 1879

Eniclases Waterhouse, 1879: 66 .

Type species. Lycus luteolus Waterhouse, 1878 (by original designation).

= Trichalolus Pic, 1923: 36, hors texte; Bocak & Bocakova, 1991.

Type species. T. apertus Pic, 1923: 36 (by monotypy).

Diagnosis. Eniclases shares shortened primary costa 1 with other trichaline genera and differs in unique V-shaped pronotal carinae ( Figs 8–9 ), secondary elytral costae are always present. Male genitalia have the characteristic cup-shaped phallic apex ( Figs 11–12 ) similar to those of Schizotrichalus but unlike the latter, the phallobase has an extensive membranous sac ( Figs 11–12 ). For detailed Eniclases redescription see Bocak & Bocakova (1991).

Distribution and diversity. The highest diversity of Eniclases is known from New Guinea. Altogether, 37 formally described species are known from the island and only Eniclases moluccanus has been reported from Halmahera ( Bocak & Bocakova 1991, Bocak 2002, Bocek & Bocak 2016).

Remark. The molecular analysis supports morphology-based sister relationships of Schizotrichalus and Eniclases . Moreover, the deeply rooted Moluccan species indicate that the ancestral range of Eniclases is located in the combined area of New Guinea and the northern Moluccas.

Figures

FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm.