Berberis sect. Wallichianae, Schneider, 1905

Yu, Chih-Chieh & Chung, Kuo-Fang, 2014, Systematics of Berberis sect. Wallichianae (Berberidaceae) of Taiwan and Luzon with description of three new species, B. schaaliae, B. ravenii, and B. pengii, Phytotaxa 184 (2), pp. 61-99 : 73-74

Key to the species of Berberis sect. Wallichianae of Taiwan and Luzon

1. Leaf veins simple (simple-veined leaf), primary vein prominent with or without observable secondary veins .............................. 2

– Leaf veins complex (complex-veined leaf), secondary veins observable with reticulated tertiary veins .........................................7

2. Adaxial surface of leaves dull ..................................................................................................................................... 4. B. hayatana

– Adaxial surface of leaves shiny with/without waxy luster ................................................................................................................3

3. Fruits estylose (style absent or indistinct less than 0.5 mm) .............................................................................................................4

– Fruits stylose (style present) ..............................................................................................................................................................5

4. Flowers in a long-pediceled fascicle, pedicel 0.5–1.3 cm, sepals 4 or 5 whorls ......................................................2. B. brevisepala

– Flowers in a dense, congested fascicle, pedicel 0.2–0.3 cm, sepals 3 whorls ........................................................... 7. B. nantoensis

5. Flowers in a congested fascicle, pedicels 0.3–1.2 cm, the outer sepals narrowly-triangular, or rarely linear ..........5. B. kawakamii

– Flowers in a long-pediceled fascicle, pedicels 1.2–1.3 cm, the outer sepals or (for flowers with 5-whorled sepals) the 4-whorled sepals ovate, obovate .........................................................................................................................................................................6 6. Leaves greyish-green abaxially, spinules 1–8 along leaf margins, 4–10 mm apart, ovules 6 or 8 ......................... 11. B. tarokoensis

– Leaves green to pale green abaxially, spinules 4–12 along leaf margins, 2–6 mm apart, ovules 4 ........................12. B. barandana