Amarotypus takaheensis Larochelle and Larivière, 2022

Larochelle & Larivière, Marie-Claude, 2022, Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2022 (942), pp. 1-30 : 7

Amarotypus takaheensis Larochelle and Larivière , new species

Fig. 12 , 23 , 45

Amarotypus takaheensis Larochelle and Larivière , new species. Holotype: male (NZAC) labeled “ NEW ZEALAND FD Takahe V [=Valley] 7 Nov 2012 – 12 March 2013, 1220m, R. Leschen, A. Schnurpfeil (typed) / Chionochloa teretifolia -45.2735475. 167.62.7021[= 167.627021E]. Pit trap # (typed) / T1-15 ug (hand-written) / HOLOTYPE ♂ Amarotypus takaheensis Larochelle & Larivière, 2022 (red label; typed).” Paratype: one female (AMNZ) from Hump Mountain [=The Hump, FD], bearing a blue paratype label.

Description. Body length 5.7 mm; ovate. Head, pronotum, and elytra piceous black; abdomen rufopiceous; antennal segments 1–4 rufotestaceous, 5–11 piceous black; palpi and legs rufotestaceous. Microsculpture strong and isodiametric (almost granulate) on head, weak and isodiametric on pronotum, obsolete and isodiametric on elytra. Iridescence absent. Shiny on head, pronotum, and elytra. Metallic luster absent. Head. Labrum moderately transverse. Eyes slightly convex. Thorax. Pronotum moderately convex, obsoletely punctate and unwrinkled across base, trapezoid, widest basally; apex moderately emarginate; anterolateral angles poorly developed, angulate; anterior bead complete; sides moderately rounded, moderately sinuate posteriorly; lateral beads narrow, slightly widened from apex to base; lateral depressions absent anteriorly, wide posteriorly; posterolateral angles acute, moderately projected laterally; laterobasal foveae with outer fovea oblong, narrow, shallow, and inner fovea oblong, moderately wide, moderately deep; posterior bead obsolete. Legs. Pro- and mesotarsomeres 1–4 of both sexes moderately dilated. Elytra. Ovate. Fused along suture (hindwings vestigial). Strongly convex. Shoulders slightly obtuse. Sides strongly rounded. Striae deep, coarsely punctate; stria 3 with three poorly developed, fine setiferous punctures. Scutellar striole very long (about two-thirds of elytral length). Intervals slightly convex. Umbilicate series with 15 setiferous punctures. Apices obtusely rounded. Aedeagus. Lateral view ( Fig. 12 ): strongly arcuate, slender, moderately widened in apical half; base slightly convex dorsally; middle strongly convex dorsally, strongly concave ventrally, without dorsal membranous area; apex moderately concave dorsally and ventrally, with extreme tip narrow, rounded, hooked dorsally. Dorsal view: asymmetrical (ostium of membranous area deflected to the right).

Material examined. 2 specimens ( AMNZ, NZAC).

Geographic distribution ( Fig. 45 ). South Island: FD –Fiordland National Park (Murchison Mountains, Takahe Valley; The Hump).

Ecology. Alpine. Epigean. Tussock grassland. Open ground. Probably nocturnal.

Biology. Seasonality: January, March. Predacious (based on mouthpart morphology).

Dispersal power. Subapterous (incapable of flight).

Collecting technique. Pitfall trapping.

Remarks. This species is named after the type locality Takahe Valley (FD) and the Latin suffix – ensis, denoting a place, locality, or country. Amarotypus takaheensis is morphologically close to A. glasgowensis . In addition to diagnostic characters of the male genitalia, Amarotypus takaheensis has the following distinguishing features: dorsal body piceous black; antennal segments 1–4 rufotestaceous, 5–11 piceous black; elytra with isodiametric microsculpture, striae deep and coarsely punctate, stria 3 with three setiferous punctures. The two species are allopatric: A. takaheensis occurs in the southwestern South Island (FD), while A. glasgowensis is known only from the northwestern South Island (NN).

Figures

Figures 1–14. Amarotypini morphology. 1–4) Microsculpture. 1) Granulate. 2) Isodiametric. 3) Moderately transverse. 4) Very transverse. 5–7) Head. 5) Setiferous punctures medial, close to eye, frontal furrows (ff) subparallel posteriorly. 6) Punctures medial, distant from eye, furrows subparallel posteriorly. 7) Punctures postmedial, distant from eye, furrows convergent posteriorly. 8–10) Gonocoxites 1 (gc 1) and 2 (gc 2). 8) Amarotypus. 9) Amaroxenus. 10) Amarophilus. 11–14) Aedeagus, lateral. 11) Amarotypus edwardsii. 12) Amarotypus takaheensis new species. 13) Amarotypus fiordlandensis new species. 14) Amarotypus murchisonorum new species.

Figures 21–24. Dorsal habitus. 21) Amarotypus edwardsii. 22) Amarotypus simoninensis new species (holotype). 23) Amarotypus takaheensis new species (paratype). 24) Amarotypus glasgowensis new species (holotype). Scale line = 1 mm.

Figures 42–47. Species distribution maps. 42) Amarotypus glasgowensis new species. 43) Amarotypus murchisonorum new species. 44) Amarotypus simoninensis new species. 45) Amarotypus takaheensis new species. 46) Amaroxenus arnaudensis new species. 47) Amaroxenus glacialis new species.