Amaroxenus Larochelle and Larivière, 2022

Larochelle & Larivière, Marie-Claude, 2022, Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2022 (942), pp. 1-30 : 10

Genus Amaroxenus Larochelle and Larivière , new genus

Fig. 9 , 15–17 , 27–30 , 46–49

Type species. Amaroxenus kahurangiensis new species, by present designation.

Description. Body length 5.3–6.8 mm; slender; pronotum cordate or subcordate, elytra subovate; forebody narrower than elytra. Color dark, rufopiceous (brown in A. glacialis ). Metallic luster absent. Dorsal surface mostly glabrous. Head. Mandibles moderately long and curved. Eyes slightly convex (subdepressed in A. glacialis ); setiferous puncture inserted postmedially on inner side of each eye, distant from it. Frontal furrows very deep, convergent posteriorly ( Fig. 7 ). Submentum with six setae. Thorax. Pronotum moderately wide, cordate or subcordate. Laterobasal foveae single. Pronotal base narrower than elytral base. Legs. Very long. Pro- and mesotarsomeres 1–4 of both sexes moderately dilated (strongly dilated in A. glacialis ). Elytra. Subovate. Fused along suture (hindwings vestigial). Shoulders poorly developed, rounded. Striae present, consisting of impressed lines; stria 3 with two or three setiferous punctures. Scutellar striole very long (about of two-thirds of elytral length). Intervals slightly convex (depressed in A. glacialis ). Umbilicate series with 12 to 16 setiferous punctures. Aedeagus. Lateral view ( Fig. 15–17 ): strongly arcuate, stout, multistriate in apical half (not multistriate in A. glacialis ). Dorsal view: asymmetrical (ostium of membranous area deflected to the right). Parameres: right paramere with a ventral fringe of short setae (long setae in A. huttensis ). Gonocoxites. Right and left pair close to each other; gonocoxite 1 slender, subparallel, without rows of spines; gonocoxite 2 with a single subapical seta ( Fig. 9 ).

Remarks. The generic name is masculine and derived from Amarotypus (type genus of the tribe Amarotypini ) and the Greek noun xenos (stranger), alluding to differences in body shape between Amaroxenus and Amarotypus . The morphological characters unifying species of Amaroxenus are: body slender; pronotum cordate or subcordate; elytra subovate; legs very long; setiferous puncture of head inserted postmedially on inner side of each eye; frontal furrows very deep, convergent posteriorly; aedeagus (in lateral view) stout, multistriate in apical half (not multistriate in A. glacialis ); right and left pair of gonocoxites close to each other, gonocoxite 1 subparallel. The species of this genus occur throughout the South Island, in subalpine or alpine tussock grasslands and fellfields.

Figures

Figures 1–14. Amarotypini morphology. 1–4) Microsculpture. 1) Granulate. 2) Isodiametric. 3) Moderately transverse. 4) Very transverse. 5–7) Head. 5) Setiferous punctures medial, close to eye, frontal furrows (ff) subparallel posteriorly. 6) Punctures medial, distant from eye, furrows subparallel posteriorly. 7) Punctures postmedial, distant from eye, furrows convergent posteriorly. 8–10) Gonocoxites 1 (gc 1) and 2 (gc 2). 8) Amarotypus. 9) Amaroxenus. 10) Amarophilus. 11–14) Aedeagus, lateral. 11) Amarotypus edwardsii. 12) Amarotypus takaheensis new species. 13) Amarotypus fiordlandensis new species. 14) Amarotypus murchisonorum new species.

Figures 15–20. Aedeagus, lateral. 15) Amaroxenus glacialis new species. 16) Amaroxenus huttensis new species. 17) Amaroxenus kahurangiensis new species. 18) Amarophilus otagoensis new species. 19) Amarophilus lomondensis new species. 20) Amarophilus wanakensis new species.

Figures 25–28. Dorsal habitus. 25) Amarotypus fiordlandensis new species. 26) Amarotypus murchisonorum new species. 27) Amaroxenus glacialis new species. 28) Amaroxenus arnaudensis new species (holotype). Scale line = 1 mm.

Figures 29–32. Dorsal habitus. 29) Amaroxenus huttensis new species. 30) Amaroxenus kahurangiensis new species. 31) Amarophilus rotundicollis new species (holotype). 32) Amarophilus otagoensis new species. Scale line = 1 mm.

Figures 42–47. Species distribution maps. 42) Amarotypus glasgowensis new species. 43) Amarotypus murchisonorum new species. 44) Amarotypus simoninensis new species. 45) Amarotypus takaheensis new species. 46) Amaroxenus arnaudensis new species. 47) Amaroxenus glacialis new species.

Figures 48–49. Species distribution maps. 48) Amaroxenus huttensis new species. 49) Amaroxenus kahurangiensis new species.