Amarotypus murchisonorum Larochelle and Larivière, 2022

Larochelle & Larivière, Marie-Claude, 2022, Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2022 (942), pp. 1-30 : 9-10

Amarotypus murchisonorum Larochelle and Larivière , new species

Fig. 14 , 26 , 43

Amarotypus murchisonorum Larochelle and Larivière , new species. Holotype: male (NZAC) labeled “ NEW ZEALAND FD Murchison Mtns [= Mountains ] Takahe Vly [= Valley] 2013“/”14 (typed) / Pitfall trap R Leschen and others (typed) / T8 + So15ug (hand-written) / HOLOTYPE ♂ Amarotypus murchisonorum Larochelle & Larivière, 2022 (red label; typed).” Paratypes: three females (NZAC) from the same locality as the holotype, bearing blue paratype labels.

Description. Body length 6.6–6.8 mm; subovate. Head, pronotum, elytra, and abdomen rufopiceous; antennae, palpi, and tarsi rufotestaceous; femora and tibiae rufopiceous. Microsculpture strong, isodiametric (almost granulate) on head, isodiametric on pronotum, and moderately transverse (brick-like) on elytra. Iridescence present on elytra. Shiny on head, pronotum, and elytra. Metallic luster absent. Head. Labrum moderately transverse. Eyes moderately convex. Thorax. Pronotum moderately convex, finely punctate on disc, strongly wrinkled on disc and across base, subrectangular, widest about middle; apex moderately emarginate; anterolateral angles well developed, angulate; anterior bead complete; sides strongly rounded, slightly sinuate posteriorly; lateral beads narrow, slightly widened from apex to base; lateral depressions absent anteriorly, wide posteriorly; posterolateral angles obtuse; laterobasal foveae with outer fovea oblong, narrow, shallow, and inner fovea oblong, moderately wide, very deep; posterior bead obsolete. Legs. Pro- and mesotarsomeres 1–4 of both sexes strongly dilated. Elytra. Oblong. Fused along suture (hindwings vestigial). Moderately convex. Shoulders slightly obtuse. Sides moderately rounded. Striae shallow, finely punctate; stria 3 with four poorly developed, fine setiferous punctures. Scutellar striole very long (about two-thirds of elytral length). Intervals slightly convex. Umbilicate series with 15 setiferous punctures. Apices obtusely rounded. Aedeagus. Lateral view ( Fig. 14 ): moderately arcuate, slender, strongly widened apically; base slightly convex dorsally; middle moderately convex dorsally, moderately concave ventrally, without dorsal membranous area; apex hammer-shaped, moderately biconcave dorsally, moderately concave ventrally, with extreme tip very wide, square, curved downward. Dorsal view: asymmetrical (ostium of membranous area deflected to the right).

Material examined. 135 specimens ( NZAC).

Geographic distribution ( Fig. 43 ). South Island: FD –Fiordland National Park (Murchison Mountains, Takahe Valley).

Ecology. Subalpine. Arboreal. Wet forests (beech). Shaded ground. Probably nocturnal.

Biology. Seasonality: November–March. Predacious (based on mouthpart morphology).

Dispersal power. Subapterous (incapable of flight). Moderate runner.

Collecting technique. Pitfall trapping.

Remarks. This species is named after the Murchison Mountains (FD) where the type locality Takahe Valley is situated and the Latin suffix – orum, meaning ‘of ‘ (of the Murchison Mountains). Amarotypus murchisonorum is morphologically close to A. fiordlandensis . In addition to diagnostic characters of the male genitalia, Amarotypus murchisonorum has the following distinguishing features: body larger, length 6.6–6.8; metallic luster absent; antennae rufotestaceous; pronotum moderately convex, sides slightly sinuate posteriorly, posterolateral angles obtuse; elytra oblong, striae shallow and finely punctate, stria 3 with four setiferous punctures. Amarotypus murchisonorum is known only from the type locality.

Figures

Figures 1–14. Amarotypini morphology. 1–4) Microsculpture. 1) Granulate. 2) Isodiametric. 3) Moderately transverse. 4) Very transverse. 5–7) Head. 5) Setiferous punctures medial, close to eye, frontal furrows (ff) subparallel posteriorly. 6) Punctures medial, distant from eye, furrows subparallel posteriorly. 7) Punctures postmedial, distant from eye, furrows convergent posteriorly. 8–10) Gonocoxites 1 (gc 1) and 2 (gc 2). 8) Amarotypus. 9) Amaroxenus. 10) Amarophilus. 11–14) Aedeagus, lateral. 11) Amarotypus edwardsii. 12) Amarotypus takaheensis new species. 13) Amarotypus fiordlandensis new species. 14) Amarotypus murchisonorum new species.

Figures 25–28. Dorsal habitus. 25) Amarotypus fiordlandensis new species. 26) Amarotypus murchisonorum new species. 27) Amaroxenus glacialis new species. 28) Amaroxenus arnaudensis new species (holotype). Scale line = 1 mm.

Figures 42–47. Species distribution maps. 42) Amarotypus glasgowensis new species. 43) Amarotypus murchisonorum new species. 44) Amarotypus simoninensis new species. 45) Amarotypus takaheensis new species. 46) Amaroxenus arnaudensis new species. 47) Amaroxenus glacialis new species.