Amaroxenus glacialis Larochelle and Larivière, 2022

Larochelle & Larivière, Marie-Claude, 2022, Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2022 (942), pp. 1-30 : 11-12

Amaroxenus glacialis Larochelle and Larivière , new species

Fig. 15 , 27 , 47

Amaroxenus glacialis Larochelle and Larivière , new species. Holotype: male (NZAC) labeled “ NEW ZEALAND WD Chancellor area Fox Glacier 30-Jan-05 (typed) / Under rock at c 1400m (typed) / Donated by J. Nunn (typed) / HOLOTYPE ♂ Amaroxenus glacialis Larochelle & Larivière, 2022 (red label; typed).” Paratypes: one female (NZAC) from the same locality as the holotype and one male (AMNZ) from Mount Moltke , bearing blue paratype labels.

Description. Body length 6.5–6.8 mm. Head, pronotum, elytra, and abdomen brown; antennal segments 1–4 rufotestaceous, 5–11 black; palpi rufotestaceous; femora and tibiae rufopiceous; tarsi rufotestaceous. Microsculpture strong (weak in other species), isodiametric on head (almost granulate), pronotum, and elytra. Iridescence absent. Dull on head, pronotum, and elytra. Head. Labrum strongly transverse. Eyes subdepressed (slightly convex in other species). Thorax. Pronotum moderately convex, obsoletely punctate in laterobasal foveae, obsoletely wrinkled on disc and across base, subcordate, moderately wide (more so than in other species), widest before middle; apex subtruncate; anterolateral angles poorly developed, obtuse; anterior bead incomplete, obsolete laterally (near posterolateral angles); sides moderately rounded, not sinuate posteriorly; lateral beads narrow throughout; lateral depressions absent anteriorly, narrow posteriorly; posterolateral angles rectangular; laterobasal foveae rounded, very wide, very deep, reaching lateral depressions; posterior bead incomplete, obsolete laterally. Legs. Pro- and mesotarsomeres 1–4 of both sexes strongly dilated. Elytra. Moderately convex. Sides strongly rounded. Striae deep, coarsely punctate; stria 3 with two poorly developed, fine setiferous punctures. Intervals depressed. Umbilicate series with 12 to 14 setiferous punctures. Apices angulate. Aedeagus. Lateral view ( Fig. 15 ): strongly arcuate, stout, strongly widened and not multistriate in apical half; base slightly convex dorsally; middle strongly convex dorsally, strongly concave ventrally, with dorsal membranous area narrow and long in apical half; apex slightly concave dorsally and ventrally, with extreme tip very wide, rounded. Dorsal view: asymmetrical (ostium of membranous area deflected to the right).

Material examined. 3 specimens ( AMNZ, NZAC).

Geographic distribution ( Fig. 47 ). South Island: WD –Westland National Park (Fox Glacier, Chancellor area; Mount Moltke).

Ecology. Subalpine. Epigean. Moraine and tussock area. Open ground. Nocturnal; hides during the day under stones.

Biology. Seasonality: January. Predacious (based on mouthpart morphology).

Dispersal power. Subapterous (incapable of flight).

Collecting technique. Turning stones.

Remarks. This species is named after the type locality Fox Glacier (WD). Amaroxenus glacialis is morphologically close to A. arnaudensis and can be distinguished from it by the following features: dorsal body dull brown; microsculpture strong; eyes subdepressed; pro- and mesotarsomeres 1–4 of both sexes strongly dilated; pronotum moderately convex, with posterolateral angles rectangular and laterobasal foveae very wide, very deep, reaching lateral depressions; elytra moderately convex, striae deep and coarsely punctate, intervals depressed. The two species are allopatric: A. glacialis occurs in the Westland region (WD) of the South Island, while A. arnaudensis is known only from the Marlborough region (MB) in the northeast of the South Island.

Figures

Figures 15–20. Aedeagus, lateral. 15) Amaroxenus glacialis new species. 16) Amaroxenus huttensis new species. 17) Amaroxenus kahurangiensis new species. 18) Amarophilus otagoensis new species. 19) Amarophilus lomondensis new species. 20) Amarophilus wanakensis new species.

Figures 25–28. Dorsal habitus. 25) Amarotypus fiordlandensis new species. 26) Amarotypus murchisonorum new species. 27) Amaroxenus glacialis new species. 28) Amaroxenus arnaudensis new species (holotype). Scale line = 1 mm.

Figures 42–47. Species distribution maps. 42) Amarotypus glasgowensis new species. 43) Amarotypus murchisonorum new species. 44) Amarotypus simoninensis new species. 45) Amarotypus takaheensis new species. 46) Amaroxenus arnaudensis new species. 47) Amaroxenus glacialis new species.