Amarotypus Bates, 1872

Larochelle & Larivière, Marie-Claude, 2022, Synopsis of the tribe Amarotypini in New Zealand (Coleoptera: Carabidae), Insecta Mundi 2022 (942), pp. 1-30 : 4

Genus Amarotypus Bates, 1872

Fig. 8, 11–14 , 21–26 , 40–45

Amarotypus Bates, 1872: 50 .

Type species. Amarotypus edwardsii Bates, 1872 , by monotypy.

Description. Body length 4.6–7.0 mm; stout, ovate or subovate; forebody as wide as elytra. Color dark, rufopiceous, rufotestaceous or piceous black. Metallic luster present (aeneous) or absent. Dorsal surface mostly glabrous. Head. Mandibles short, strongly curved. Eyes slightly or moderately convex; setiferous puncture inserted medially on inner side of each eye, close to it. Frontal furrows shallow, subparallel posteriorly ( Fig. 5 ). Submentum with four setae. Thorax. Pronotum very wide, trapezoid or subrectangular. Laterobasal foveae double. Pronotal base as wide as elytral base. Legs. Moderately long. Pro- and mesotarsomeres 1–4 of both sexes moderately dilated (strongly dilated in A. murchisonorum ). Elytra. Oblong or ovate. Fused along suture (hindwings poorly developed, vestigial) or free along suture (hindwings more developed, 65–75% of elytral length in A. edwardsii ). Shoulders well developed, obtuse. Striae present, consisting of impressed lines, or replaced by rows of punctures; stria 3 or row 3 with one to four setiferous punctures. Scutellar striole very long, about two-thirds of elytral length (moderately long, about a quarter of elytral length in A. simoninensis ). Intervals slightly convex (subdepressed in A. edwardsii ). Umbilicate series with 13–15 setiferous punctures. Aedeagus. Lateral view ( Fig. 11–14 ): slightly to strongly arcuate, slender (stout in A. edwardsii ), not multistriate in apical half. Dorsal view: asymmetrical (ostium of membranous area deflected to the left or to the right). Parameres: right paramere with a ventral fringe of long setae. Gonocoxites. Right and left pair close to each other; gonocoxite 1 stout, with rows of spines, gonocoxite 2 with two subapical setae ( Fig. 8 ).

References. Larochelle and Larivière 2001: 40 (catalogue), 2007: 29 (description, ecology, geographic distribution, references).

Remarks. The morphological characters unifying species of Amarotypus are: setiferous puncture of head inserted medially on inner side of each eye, close to it; frontal furrows shallow, subparallel posteriorly; submentum with four setae; laterobasal foveae of pronotum double; right and left pair of gonocoxites close to each other, gonocoxite 1 stout, with rows of spines, gonocoxite 2 with two subapical setae. Members of this genus usually occur in lowland to subalpine forests of the North, South, and Stewart Islands. Two species have been found in alpine tussock grasslands.

Figures

Figures 1–14. Amarotypini morphology. 1–4) Microsculpture. 1) Granulate. 2) Isodiametric. 3) Moderately transverse. 4) Very transverse. 5–7) Head. 5) Setiferous punctures medial, close to eye, frontal furrows (ff) subparallel posteriorly. 6) Punctures medial, distant from eye, furrows subparallel posteriorly. 7) Punctures postmedial, distant from eye, furrows convergent posteriorly. 8–10) Gonocoxites 1 (gc 1) and 2 (gc 2). 8) Amarotypus. 9) Amaroxenus. 10) Amarophilus. 11–14) Aedeagus, lateral. 11) Amarotypus edwardsii. 12) Amarotypus takaheensis new species. 13) Amarotypus fiordlandensis new species. 14) Amarotypus murchisonorum new species.

Figures 21–24. Dorsal habitus. 21) Amarotypus edwardsii. 22) Amarotypus simoninensis new species (holotype). 23) Amarotypus takaheensis new species (paratype). 24) Amarotypus glasgowensis new species (holotype). Scale line = 1 mm.

Figures 25–28. Dorsal habitus. 25) Amarotypus fiordlandensis new species. 26) Amarotypus murchisonorum new species. 27) Amaroxenus glacialis new species. 28) Amaroxenus arnaudensis new species (holotype). Scale line = 1 mm.

Figures 36–41. Species distribution maps. 36) Amarophilus lomondensis new species. 37) Amarophilus otagoensis new species. 38) Amarophilus rotundicollis new species. 39) Amarophilus wanakensis new species. 40) Amarotypus edwardsii. 41) Amarotypus fiordlandensis new species.

Figures 42–47. Species distribution maps. 42) Amarotypus glasgowensis new species. 43) Amarotypus murchisonorum new species. 44) Amarotypus simoninensis new species. 45) Amarotypus takaheensis new species. 46) Amaroxenus arnaudensis new species. 47) Amaroxenus glacialis new species.