Enicocephalidae, Stal, 1860

Kment, Petr, Baňař, Petr, Bílý, Svatopluk, Pluot-Sigwalt, Dominique, Polhemus, Dan A. & Schuh, Randall T., 2019, In memoriam of Professor Pavel Štys (1933 - 2018): biography, memories, bibliography and list of described taxa, Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (2), pp. 351-379 : 371-373

Family Enicocephalidae

Alienates thomasi Baňař & Štys, 2015 in BAŇAŘ et al. (2015a: 392) . Holotype: ♀, Venezuela: Aragua Province, Rancho Grande (= Henry Pittier National Park) (USNM). Current status: Valid taxon.

Alkowdiella Štys, 2002 a: 326. Type species: Alkowdiella raunoi Štys, 2002 . Gender: feminine (original designation). Current status: Valid taxon (see AUKEMA et al. 2013).

Alkowdiella paleceki Štys, 2002 a: 333. Holotype: ♂, Yemen: ‘ Al Kowd’ (RLRF → NMWC).Current status:Valid taxon (seeAUKEMA et al.2013).

Alkowdiella raunoi Štys, 2002 a: 330. Holotype: ♂, Yemen: ‘ Al Kowd’ (RLRF → NMWC).Current status:Valid taxon (seeAUKEMA et al.2013).

Ciucephalus Štys, 1982c: 500 . Type species: Ciucephalus wygodzinskyi Štys, 1982 . Gender: masculine (original designation).Current status: Valid taxon (see ŠTYS 2002b).

Ciucephalus wygodzinskyi Štys, 1982c: 500 . Holotype: ♂, New Caledonia: ʻCiu, nr. Mt. Canalaʼ (AMNH). Current status: Valid taxon.

† Disphaerocephalinae Štys, 1969d:355 . Type genus: Disphaerocephalus Cockerell, 1917 . Current status: Valid taxon.

Embolorrhinus (Ceratotrachelus) villiersi Štys, 1969 a: 282. Holotype: ♀, Guinea: ‘ Nimba , Ziela’ (MNHN). Current status: Valid taxon (see VILLIERS 1969).

Gourlayocoris Štys, 2002b: 340 . Type species: Phthirocoris mirabilis Gourlay, 1952 . Gender: masculine (original designation). Current status: Valid taxon (see LARIVIÈRE & LAROCHELLE 2004).

Heissaptera Štys & Baňař, 2006b:683 . Type species: Heissaptera janaki Štys & Baňař, 2006 .Gender:feminine (original designation). Current status: Valid taxon.

Heissaptera janaki Štys & Baňař, 2006b: 684 . Holotype: ♀ (apterous), Mauritius: NW Mauritius, Le Pouce Mt. , 20°11′48″S 57°31′21″E (NMPC, see KMENT et al. 2015: 437). Current status: Valid taxon.

Henicocorinus Štys, 1988 a: 159. New substitute name for Henicocoris Villiers, 1969 , preoccupied by Henicocoris Woodward, 1968 ( Heteroptera : Henicocoridae ). Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b).

Henschiella (Trarza) congoensis Štys, 1968: 333 , 340. Holotype: sex unknown – probably ♀, Democratic Republic of the Congo: ‘Congo: Katanga, Kipushi’ (BMNH). Current status: Valid taxon (see VILLIERS 1969).

Hoplitocoris (Hoplitocoris) ethiopicus Štys, 1969f: 75 . Holotype: ♀, Ethiopia: ‘ Addis Ababa [sic!], “ A.O.I. Scioa ” ’ (BMNH). Current status: Valid taxon (see VILLIERS 1969).

Hoplitocoris (Iphlotocoris) isaroginus Štys, 1981 a: 93. Holotype: ♂, Philippines: Luzon, Prov. Camarines Sur, Mt.Isarog , 20 km E of Naga (13.36 N, 132.12 E)’ (BPBM). Current status: Valid taxon.

Hoplitocoris (Iphlotocoris) xandrae Štys, 1981 a: 97. Holotype: ♀, Philippines: ‘ Negros I., Dumaguete (9.20 N, 123.18 E), Camp Lookout’ (AMNH). Current status: Valid taxon.

Iphlotocoris Štys, 1981a: 90 (as subgenus of Hoplitocoris Jeannel, 1942 ). Type species: Hoplitocoris isaroginus Štys, 1981 . Gender:masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b).

Kulichoderes Štys, 2002 a: 322. Type species: Systelloderes iranicus Štys, 1970 .Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b, AUKEMA et al. 2013).

Megenicocephalinae Štys, 1989a: 75. Type genus: Megenicocephalus Usinger, 1945 . Current status: Valid taxon (see KERZHNER 1995a, ŠTYS 1995a).

Monteithostolinae Štys, 1981b: 412. Type genus: Monteithostolus Štys, 1981 . Current status:Valid taxon, downgraded to tribe of Phthirocorinae Jeannel, 1942 (ŠTYS 1989a: 74; see ŠTYS 1995a).

Monteithostolus Štys, 1981b:413 . Type species: Monteithostolus genitalis Štys, 1981 . Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b).

Monteithostolus caledonicus Štys, 1982c:496 . Holotype: ♀, New Caledonia:ʻKavatch, near Hienghéneʼ (NZAC). Current status: Valid taxon.

Monteithostolus genitalis Štys, 1981b: 413 . Holotype:♂, New Caledonia: ‘ Houailou River , 300′’ (UQIC). Current status: Valid taxon.

Neoncylocotis kotejai Štys & Baňař, 2006 a: 170. Holotype:♂, Ecuador: ‘ Pastaza Province, 2 km W of Santa Clara (01.18 S, 77.52 W)’ (QCAZ, currently deposited in MMBC). Current status: Valid taxon.

Oncylocotis cupressi Štys, 1999 a: 207. Holotype: ♂ (submicropterous), Tanzania: Arusha, ‘S slope of Mt. Hanang (gorge of Himit; 4.27 S, 35.24 E)’ (MRAC). Current status: Valid taxon, Oncylocotis (Oncylocotis) cupressi Štys, 1999 (see ŠTYS 2002b: 344).

Oncylocotis froeschneri Štys, 1986b: 149 . Holotype: ♂ (extremely brachypterous), Democratic Republic of the Congo: ʻZaire, Kivu, Uviraʼ (MRAC). Current status: Valid taxon, Oncylocotis (Oncylocotis) froeschneri Štys, 1986 (see ŠTYS 2002b: 344).

Oncylocotis (Oncylocotis) inexpectatus Štys, Baňař & Drescher, 2010: 29 . Holotype: ♀, Malaysia: Sabah, Poring Hot Springs , 6°02′41.78″N, 116°42′01.97″E (MMBC, deposited in alcohol). Current status: Valid taxon.

Oncylocotis lautereri Baňař & Štys, 2013: 322 . Holotype: ♀, Australia: Northern Territory, Litchfield N.P., Tjaynera Falls , 13°15′S 130°44′E (MAGNT). Current status: Valid taxon.

Oncylocotis neotenicus Štys, 1982b: 486 . Holotype: ♂ (micropterous), Indonesia: ʻSumatraʼ (RMNH). Current status: Valid taxon, Oncylocotis (Oncylocotis) neotenicus Štys, 1982 (see ŠTYS 2002b: 344).

Oncylocotis occipitalis meridionalis Štys, 1969 a:281. Holotype: ♀, South Africa:‘S. Africa, Natal, Bell Morley’ (BMNH). Current status:Junior subjective synonym of Oncylocotis (Oncylocotis) occipitalis (Jeannel, 1942) ( VILLIERS 1969: 66; see KERZHNER 1995a).

Oncylocotis reductus Štys, 1999 a: 205. Holotype: ♂ (submicropterous), Tanzania: Arusha, ‘E slope of Mt. Oldeani’ (MRAC). Current status: Valid taxon, Oncylocotis (Oncylocotis) reductus Štys, 1999 (see ŠTYS 2002b: 344).

Oncylocotis stehliki Baňař & Štys, 2013: 318 . Holotype: ♂, Australia: Northern Territory, Nitmiluk N.P., 24°18′S 131°34′E, Edit Falls (MAGNT). Current status: Valid taxon (see BAŇAŘ & ŠTYS 2011).

Owenoderes Štys, 2002 a:323. Type species: Systelloderes oculatus Štys, 1969 .Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b).

Phaenicocleus Štys & Baňař, 2009: 22 . Type species: Phaenicocleus sabahensis Štys & Baňař, 2009 .Gender:masculine (original designation). Current status: Valid taxon.

Phaenicocleus granulosus Baňař & Štys, 2011:17 . Holotype: ♀, Malaysia: Sabah, Tambunan distr., Crocker Range , 1560–1650 m, near the pass (KotaKinabalu–Tambunan road) (MHNG).Current status:Valid taxon.

Phaenicocleus minor Štys & Baňař, 2009: 24 . Holotype: ♂, Malaysia: Sabah, Tambunan distr., Crocker Range, 51–52 km of road Kota Kinabalu–Tambunan (MHNG, preserved in alcohol). Current status: Valid taxon.

Phaenicocleus sabahensis Štys & Baňař, 2009: 28 . Holotype: ♂, Malaysia: Sabah, Tambunan distr., Crocker Range , 1560–1650 m, near the pass (Kota Kinabalu–Tambunan road) (MHNG). Current status: Valid taxon.

Phaenicocleus schwendingeri Štys & Baňař, 2009: 30 . Holotype: ♂, Malaysia: Sabah, West Coast Residency, Kinabalu Park, Mt. Kinabalu , 2600 m, near Layang-Layang (MHNG, preserved in alcohol). Current status: Valid taxon.

Phallopiratinae Štys, 1985c: 271. Type genus: Phallopirates Štys, 1985 . Current status:Valid taxon (see ŠTYS 1989a, WYGODZINSKY & SCHMIDT 1991, KERZHNER 1995a, ŠTYS 1995a).

Phallopirates Štys, 1985c:271 . Type species: Phallopirates palawanicus Štys, 1985 . Gender: masculine (original designation).Current status: Valid taxon (see ŠTYS 2002b).

Phallopirates borneensis Štys, 1985c: 295 . Holotype: ♂, Malaysia: ʻBorneo, Sarawak, 4th div. G. Mulu N. P., Karangas Forest , plot 3rd Campsʼ (BMNH). Current status: Valid taxon.

Phallopirates culionicus Štys, 1985c: 291 . Holotype: ♂, Philippines: ʻCulion I., 6 km W. Culionʼ (BPBM). Current status: Valid taxon.

Phallopirates malayicus Štys, 1985c:286 . Holotype:♂, Malaysia:ʻJohore St. (»Johar«), Layang Layangʼ (AMNH).Current status: Valid taxon.

Phallopirates palawanicus Štys,1985c:272 . Holotype:♂, Philippines:ʻPalawan Is., Mantalingajan , Pinigisanʼ (ZMUC).Current status:Valid taxon.

Phthirocorisella Štys, 1986 a: 63. Type species: Phthirocorisella petrophila Štys, 1986 . Gender: feminine (original designation). Current status: Valid taxon (see ŠTYS 2002b).

Phthirocorisella petrophila Štys, 1986 a: 65. Holotype: ♂, Papua New Guinea: ʻEdie Creekʼ (BPBM). Current status: Valid taxon.

Phthirostenus Štys, 2002b: 341 . Type species: Phthirocoris magnus Woodward, 1956 . Gender: masculine (original designation). Current status: Valid taxon (see LARIVIÈRE & LAROCHELLE 2004).

Proboscidopirates ericguilberti Baňař & Štys, 2015 in BAŇAŘ et al. (2015b: 408) . Holotype: ♀, Madagascar: border of Andasibe N.P., Feon’ny Ala hotel surroundings (MMBC).Current status:Valid taxon.

Proboscidopirates rugulosus Baňař & Štys, 2015 in BAŇAŘ et al. (2015b: 412) . Holotype: ♀, Madagascar: Andasibe N.P.,Ambatomandondona (MMBC). Current status: Valid taxon.

Pseudohenschiella hauseri Baňař & Štys, 2006: 8 . Holotype: ♀, Madagascar: Antananarivo prov., Ankaratra hills, Manjakatompo Forest Station, near the summit of Anosiarivo (MHNG). Current status: Valid taxon.

† Pyrenicocephalus Štys, 2010: 456 . Type species: Pyrenicocephalus jarzembowskii Štys, 2010 . Gender: masculine (original designation). Current status: Valid taxon.

† Pyrenicocephalus jarzembowskii Štys, 2010: 456 . Holotype: head, Great Britain, England, Kent, Isle of Sheppey, Paddy’s Point. Type horizont: Early Eocene. Upper London Clay, Division E, unit SH-12c (BMNH: Department of Palaeontology ). Current status: Valid taxon.

Stenopirates jeanneli Štys, 1970d: 235 . Holotype: ♂, India: ‘ Sikkim, Singhik’ (BMNH).Current status: Valid taxon (see KERZHNER 1995a).

Stenopirates montanus Štys, 1970d: 231 . Holotype: ♂, India: ‘N. W. India, Kumaun , Gori Valley’ (BMNH). Current status: Valid taxon (see KERZHNER 1995a).

Stenopirates niger Štys, 1970d: 231 . Holotype: ♂, India: ‘ N. W. Bengal, Darjeeling,“Gopaldhara,BW.” ’ (BMNH).Current status:Valid taxon (see KERZHNER 1995a).

Systelloderes iranicus Štys, 1970c: 443 . Holotype: ♂, Iran: ‘ Kulichan (region of Bakhtiari and Lurs)’ (ZMAS). Current status: Valid taxon, Kulichoderes iranicus (Štys, 1970) (ŠTYS 2002a: 322; see AUKEMA et al. 2013).

Systelloderes loebli Štys & Baňař, 2007: 4 . Holotype: ♀, New Caledonia: Mount Koghi (MHNG). Current status: Valid taxon.

Systelloderes oculatus Štys, 1969 g in VILLIERS (1969: 219). Holotype:♂, Nigeria (BMNH). Note : Due to the delay in publication, the intended original description (see ŠTYS 1970c: 446) was predated by VILLIERS’ (1969) monograph (see ŠTYS 2002a: 323). The detailed type locality is Nigeria: ‘ Owena’ (ŠTYS 1970c). Current status: Valid taxon, Owenoderes oculatus (Štys, 1969) (ŠTYS 2002a: 323).

Systelloderes uvarovi Štys, 1970c: 440 . Holotype: ♂, Iraq: ‘ Kurdistan , Ugla Post on the road Mosul-Nisibin’ (BMNH). Current status: Valid taxon, Ugloderes uvarovi (Štys, 1970) (ŠTYS 2002a: 323, see AUKEMA et al. 2013).

Ugloderes Štys, 2002 a: 323. Type species: Systelloderes uvarovi Štys, 1970 .Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b, AUKEMA et al. 2013).

Utukhengal Štys, 2002a: 324. Type species: Systelloderes utukhengal Linnavuori, 1984 . Gender: masculine (original designation). Current status: Valid taxon (see ŠTYS 2002b).

Vuorillina Štys, 1986c:257. Type species: Vuorillina carayoni Štys, 1986 . Gender: feminine (original designation). Current status: Valid taxon (see KERZHNER 1995a, ŠTYS 2002b).

Vuorillina carayoni Štys, 1986c: 258 . Holotype: ♂, Saudi Arabia: ʻS., nr. Athnen (montane locality in a Juniperus zone; near Abba, 18.00 n, 42.34 E, in Asir Mts)ʼ (RLRF, NMWC, see KMENT & CARAPEZZA 2017: 536). Current status: Valid taxon (see KERZHNER 1995a).

Xenicocephalus josifovi Štys & Baňař, 2008 a: 360. Holotype: ♂, Suriname: Onverdacht , Para Distr., 5°36′N 55°09′W (RMNH). Current status: Valid taxon.

Xenicocephalus tomhenryi Štys & Baňař, 2018: 35 . Holotype: ♀, Ecuador: Pichincha Province, camino Aloag-Tandapi (USNM). Current status: Valid taxon.