Dicerogastra costigerodes ( Poole, 1989 )

Ronkay, László, Gyulai, Péter & Ronkay, Gábor, 2017, The Survey Of The Himalayan-Sino-Tibetan Species Of The Genus Dicerogastra Fletcher, 1961 (Lepidoptera: Noctuidae, Hadeninae), With The Description Of Two New Species From China, Acta Zoologica Academiae Scientiarum Hungaricae 63 (1), pp. 17-28 : 18-22

Dicerogastra costigerodes ( Poole, 1989)

( Figs 3–6 , 17–18 , 26 )

Poliacostigerodes Poole, 1989, LepidopterorumCatalogus 118 (1): 822. AnobjectiverePlace- mentnameof Agrotiscostigera Moore , 1881, necWalker, 1858. TyPelocality: [ India, HimachalPradesh] Punjab, Solun. LectotyPe: female, incoll. BMNH.

TyPematerialeXamined – LectotyPefemale , “Solun, Punjab, CaPt. Raed”, “Moore Coll., 94-106.”, “ LectotyPe ” (blueringedlabel); slideNo. HM6668 f/BMNoct. 15514f (coll. BMNH); ParalectotyPemale, “CherraPundi”, “coll. Atkinson ”, “Origin.” (Pinklabel), “Agrotiscostigera m tyPe) Moore”; “coll. Staudinger, K. 342.”, “1279”, “Miselia (Polia) costigeraMoore, Boursindet.” (coll. ZMHU) .

AdditionalmaterialeXamined – NePal. 1 male, Langtang , 1950 m, 1,5 kmNEDhunche, 24.XI.1994, leg. Csorba & Ronkay, slideNo. RL 11287m (coll. HNHM) ; 1 female, Langtang , 2850 m, 3 km SE Syabru, 85°21’E, 28°07’N, 27.IX.1994, leg. G. Csorba & L. Ronkay, slide No. RL11676 f (coll. HNHM) ; 1 male, AnnaPurnaregion , Sudame, 1250 m, 24–26.III.2000, M. Hreblay & L. Németh, slideNo. GYP 4373m (coll. PGY) ; 1 male, GaneshHimal , Kausing Danda, aboveKhurPudanda 4100 m, 20–21.IX.1995, leg. P. Gyulai & A. Garai (coll. PGY) ; 2 males, Ganesh Himal , 2 km NW of Nesim, 2300 m, 22–23.IX.1995, leg. P. Gyulai & A. Garai (coll. PGY) ; 1 male, GaneshHimal , Gadlang, 2600 m, 22–23.IX.1995, leg. P. Gyulai & A. Ga- rai (coll. PGY) ; 1 male, GaneshHimal , Mailungkhala, 1100 m, 25–26.IX.1995, leg. P. Gyulai & A. Garai (coll. PGY) ; 1 male, GaneshHimal , KhalcoPkharkha, 3400 m, 17–19.IX.1995, leg. P. Gyulai & A. Garai (coll. PGY) ; 1 male, Ganesh Himal , 2 km S of Somdang, 3000 m, 3–4. IV.1995, leg. Gy. M. László & G. Ronkay (coll. PGY) .

Diagnosis – Dicerogastracostigerodes canbedistinguishedfromallother easternAsiaticcongenersbythebroadandunicolorousochreous- beigefore- wingcostalstriPe; thecostalareaofallthreeotherrelativesisnotoronly slightlyPalerthanthemedianareaofthewing, andsuffusedwithvariably darkbrown- grey. Thewhitishsubterminallineisalmoststraightattheme- dian veins (between M 1 –M 3) in D. costigerodes and D. ferrisparsa while this sectionofthesubterminallineisangledinwardsin D. euxoides sP. n. and D. jensergabori sP. n., makingthesubterminallinesinuousinthelattertwosPe- cies. TheantemedialandPostmedialcrosslinesarelessdistinct, lessstrongly sinuousthanintheeXternallymostsimilar D. ferrisparsa andtheforewingsare somewhatnarrower, moreelongated, withmorePointedaPeX.

Themalegenitaliaof D. costigerodes ( Figs 17–18 ) differfromthoseof D. ferrisparsa ( Figs 15–16 ) bythesmallerandmoreroundedcucullussittingon longerandstraighterneck, muchlongerPin- likebasalsetaeofharPe, slen- dereramPullarProcessandtheratherS-shaPedvesicaarmedbymuchshort- ercornutifieldsituatedmedially; from D. euxoides bythelargerandmore roundedcucullushavingthinnerneck, muchshorterandthinnerbasalsetae ofharPe, finer, moreventrallyProjectingamPullarProcess, andtheconsider- ablylonger, moremediallylocatedcornutifieldonthemoreS-shaPedvesica.

ThefemalegenitaliaofthefourHimalayan- Sino- Tibetan Dicerogastra sPecies ( Figs 25–29 ) arerathersimilar, withclearlyrecognisablesPecificdifferences. ThetwosouthernHimalayansPecies, D. costigerodes and D. ferrisparsa , haveshorterantrum, somewhatthickenedPosteriorandlongitudinally ribbedanteriorPartofductusbursae, andthebasalandmedialsectionsofthe innercurveofaPPendiX (cerviX) bursaeisalsosclerotised-ribbed ( seetheFigs 25–26 ). ThesPecificdifferencesbetweenthemarefoundinthesizeandshaPe ofthesefeatures: D. costigerodes hasmoreelongatedantrum, lessdilatedPos- teriorendandshorterbutmoredenselyribbedanteriorsectionofductusbur- sae, andtheaPPendiXbursaeisonlybasallyribbed, withoutsclerotisedarea inthemedialPartofitsinnercurve.

InthetwoChinesesPecies, theantrumislonger, theductusbursaeis smoothlysclerotisedinitsentirelength, withoutlongitudinalribsatjunction tocorPusbursaebutwithastrong, sclerotisedfold, theinnercurveofaPPen- diXbursaeisgenerallymembranousbuttheaPicalsectionmaybesclerotised ( Figs 27–29 ).

Distribution – SouthernHimalayan. ThesPeciesisknownfromIndia, NePalandBhutan.

holotyPe, male ( GYP); 8 = labels of the holotyPe

holotyPe, male ( BMNH); 14 = labels of the holotyPe

holotyPe, GYP 2333m, 19 = clasPing aPParatus, 20 = aedeagus

Figures

Figs 1–8. Adults of Dicerogastra sPecies. 1–2 = Dicerogastra ferrisparsa (HamPson, 1894), 1 = lectotyPe, male (BMNH), 2 = labels of the lectotyPe; 3–4 = D. costigerodes (Poole, 1989), 3 = ParalectotyPe, male (BMNH), 4 = labels of the ParalectotyPe; 5–6 = D. costigerodes (Poole, 1989), 5 = lectotyPe, female (BMNH), 6 = labels of the lectotyPe; 7–8 = D. euxoides sP. n., 7 =

Figs 15–20. Male genitalia of Dicerogastra sPecies. 15–16 = Dicerogastra ferrisparsa (HamPson, 1894), RL11282m, 15 = clasPing aPParatus, 16 = aedeagus; 17–18 = D. costigerodes (Poole, 1989), RL11287m, 17 = clasPing aPParatus, 18 = aedeagus; 19–20 = D. euxoides sP. n.,

Figs 25–29. Female genitalia of Dicerogastra sPecies. 25 = Dicerogastra ferrisparsa (HamPson, 1894), GYP4354f; 26 = D. costigerodes (Poole, 1989), lectotyPe, BM Noct. 15514f; 27 = D. euxoides sP. n., ParatyPe, GYP1894f; 28 = D. euxoides sP. n., ParatyPe, GYP4391f; 29 = D.