Notopilo confusus, Bartlett & Lambkin, 2022

Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, Zootaxa 5220 (1), pp. 1-81 : 60-61

Notopilo confusus sp. nov.

ZooBank registration: urn:lsid:zoobank.org:act:0D9DD6A0-B7F2-462A-8FC4-592AE092857D

( Figs 21 , 57 , 129 , 172 , 185 ; Map 4)

HOLOTYPE ♁: South Australia: Adelaide, SA Feb-June 1986, S.Rondonuwu ( ANIC) . PARATYPES (16): South Australia: Adelaide, SA Feb-June 1986, S.Rondonuwu (11, ANIC) ; Mt Lofty Rgs. S. H. Curnow // Ex. Coll. S.A. Mus (1, ANIC) ; A.H. Elston Collection // Adelaide S. Australia A.H. Elston // K 304527 // 74 Opilo congruus Newm. Id by A.M. Lea (1, AM) ; Adelaide // Museum Paris Coll. Castelnau Coll. Sedillot 1935 (1, MNHN) ; Gawler S. Australia. // Pascoe Coll. 93-60. (1 ♁, NHML) ; 29959 // Nov Holl S Austr // Fry Coll. 1900 100. (1, NHML) .

Additional material. Western Australia: De Boulay // Nov.Holl. Occid. // Fry Coll. 1905-100 (2, NHML *); W. Australia Champion Bay Duboulay // Museum Paris ex Coll. R.Oberthur (1 ♁, 1, MNHN); A.H. Elston Collection // Beverley, WA // K 304535 // 74 Opilo congruus Newm. (1, AM) ; Yanchep W. Aust., Ex TuART [?] 20.xi.1970, S.J. Curry // Agriculture (Dept) Western Australia 49861 ( WADA) ; R.P. McMillan, Spencers Brook. Sept. 1948 (1, WAM E88346 ) ; WA Carlisle ( Perth ) 31°58′37″S 115°55′24″E 8 February 2000 B.P. Hanich // on outside flywire door at night (1, WAM E88389 ) .

* Note: two NHML specimens (see Fig. 185 ) were discovered to be part of the original syntype series of Opilo femoralis Westwood (see ‘comment on lectotype designation’ under Platynotum femorale stat. rev., comb. nov. below).

Diagnosis. Pronotum evenly rounded laterally, disc not heavily punctate; elytra dark with orange fasciate and apical maculations (humeral maculae faint or absent), punctation with nodules, 8 th stria beginning before fascia, at least striae 3–8 reaching apical maculae (intrastrial-punctures may become widely spaced apart towards apex); femora yellow basally, brown apically, tarsi with three ventral tarsal pads.

Description. Habitus: Fig. 172 . Total length: 7.1–9.6 mm (holotype 7.1 mm). Head ( Fig. 21 ): Cranium black, clypeus and supra-antennal elevation with a reddish or orange hue, anteclypeus, labrum, palpi and antennae orange to orange-brown; eyes separated by 0.56–0.78 eye widths (holotype 0.75); vertex slightly to moderately punctate, frons well-punctated and slightly impressed above narrowest point, punctate-rugulose below, clypeus mostly smooth with a few punctations near hypostomal suture; genae and submentum wrinkled; ratio of exterior to interior edges of terminal palpomeres about 1.8–2.2:1 (maxillae) and 2.5–3:1 (labium); antennae reaching near base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter medially-directed setae. Prothorax: Blackish to reddish-brown, pronotal arch and collar sometimes paler; pronotum 1.16–1.36 times longer than wide (holotype 1.19), sides rounded to slightly tuberculate, middle about as wide as anterior part; subapical depression vshaped, disc with deep central sulcus (sulcus smooth, more open than linear) and a poorly-defined shallow sulcus on each side, surface with small seta-associated punctations, smooth in overall appearance; short fine multi-directional setae and long erect setae. Pterothorax: Ventrites orange-brown, with short posteriorly-directed, and occasional long, setae; elytra brown with orange markings (each elytron with a large apical macula and a transverse fascia crossing elytral mid-length which is narrowed or curved at the suture, small humeral maculae uncommon); length to width ratio 2.77–3.04:1 (holotype 2.88); 8 th stria beginning between first and sixth puncture of 7 th stria, at least striae 3–8 ending at or near apical macula, punctation with small lateral nodules (most prominent anterior of fascia), punctation posterior of fascia smaller than basal punctation and may be more widely spaced, epipleurae extending into apical maculae, interstriae with erect and angled setae of slightly differing lengths (the shorter semi-decumbent setae seen in other species has become more similar to the erect setae in this species), intrafoveal setae short; hindwing with CuA 3+4 and CuA 1 cross-veins complete (CuA 1 very thin), MP 3+4 absent basad of CuA 1 cross-vein. Legs: Femora yellow basally and brown apically (apical half of profemora, apical two-fifths of mesofemora and apical third of metafemora), tibiae and tarsi brown, ventral tarsal pads yellowish; profemora only slightly thicker than other femora. Abdomen: Ventrites orange. Male genitalia: Tegmen ( Fig. 57 ) relatively broad, subparallel or wekly narrowing towards parameroid lobes, the latter tapering to a digitiform process, dorsal sinus about two-fifths tegmen length, narrow internally, wide open externally, ventral sinus one-third as long, apodeme about one-quarter tegmen length; pygidium as in Fig. 129 .

Etymology. The specific epithet confusus (Latin, meaning confounded, confused or having been brought into disorder) refers to the confusion of Westwood erroneously considering this species conspecific with Opilo femoralis Westwood.

Biology. Notopilo confusus sp. nov. has been collected from February to June (SA) and in November (WA).

Distribution (Map 4). Specimens are known from several localities between Geraldton in Western Australia and the type locality, Adelaide in South Australia.

Figures

FIGURES 12–26. (12) Notopilo variipes elytra, detail showing foveate punctures with conspicuous lateral nodules; (13) Notopilo congruus elytra, detail showing foveate punctures without lateral nodules; (14) Ancyropilus monteithi elytra, detail showing densely setose intervals; (15) Infectostriatus absentis (paratype) elytra, eighth & ninth elytral stria absent; (16) Notopilo obesus (holotype) elytra, eighth & ninth elytral stria intact; (17) Platynotum bulli (paratype), flat pronotal disc (lateral view); (18) Culcipternotus mareebensis (paratype) tarsus, with four ventral pads; (19) Notopilo reduncus tarsus, with three ventral pads; (20) Olesterus australis eye, finely-facetted; (21) Notopilo confusus eye, coarsely-facetted; (22) Monilonotum pascoei elytra, detail showing intrafoveal seta apparently absent; (23) Ancyropilus brigalowae (paratype) elytra, detail showing short intrafoveal seta; (24) Platynotum foveosetosa (holotype) elytra, detail showing very long intrafoveal seta; (25) Notopilo reduncus elytra, showing puncture rows (striae) terminating before apex; (26) Platynotum femorale elytra, showing elytral puncture rows terminating mostly near apex.

FIGURES 48–65. Tegmina, dorsal: (48–51) Notopilo, cambageicola species group—(48) Notopilo cambageicola, paratype; (49) Notopilo interfabulatus, paratype; (50) Notopilo lawnhillensis, paratype; (51) Notopilo tanybasilaris, holotype; (52) Notopilo congruus (congruus species group); (53) Notopilo reduncus (reduncus species group); (54–56) Notopilo, variipes species group—(54) Notopilo brevistriatus, paratype; (55) Notopilo gerstmeieri, holotype; (56) Notopilo variipes; (57–60) Notopilo, unplaced to species group—(57) Notopilo confusus, holotype; (58) Notopilo elstoni, holotype; (59) Notopilo katherinensis, holotype; (60) Notopilo obesus, paratype; (61) Platynotum bulli, paratype; (62) Platynotum culgoense, holotype; (63) Platynotum femorale; (64) Platynotum foveosetosa, holotype; (65) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 102–137. Male pygidiae: (102) Ancyropilus brigalowae, paratype; (103) Ancyropilus emmotti, holotype; (104) Ancyropilus exossuarius, paratype; (105) Ancyropilus monteithi, holotype; (106) Ancyropilus noonbahensis, holotype; (107) Ancyropilus packsaddlensis, holotype; (108) Ancyropilus simplex, paratype; (109) Ancyropilus tricolor, paratype; (110) Ancyropilus hornensis, paratype; (111) Culcipternotus mareebensis, paratype; (112) Infectostriatus absentis, holotype; (113) Monilonotum doddi; (114) Monilonotum pascoei; (115) Monilonotum sundholmi, paratype; (116–120) Notopilo, beswickensis species group—(116) Notopilo beswickensis, holotype; (117) Notopilo calicis, paratype; (118) Notopilo tompricensis, holotype; (119) Notopilo xanthoimprocerus, paratype; (120) Notopilo xanthoprolatus, holotype; (121–124) Notopilo, cambageicola species group—(121) Notopilo cambageicola, paratype; (122) Notopilo interfabulatus, paratype; (123) Notopilo lawnhillensis, paratype; (124) Notopilo tanybasilaris, holotype; (125) Notopilo congruus (congruus species group); (126–128) Notopilo, variipes species group—(126) Notopilo brevistriatus, paratype; (127) Notopilo gerstmeieri, holotype; (128) Notopilo variipes; (129–132) Notopilo, unplaced to species group—(129) Notopilo confusus, holotype; (130) Notopilo elstoni, holotype; (131) Notopilo katherinensis, holotype; (132) Notopilo obesus, paratype; (133) Platynotum bulli, paratype; (134) Platynotum culgoense, holotype; (135) Platynotum femorale; (136) Platynotum foveosetosa, holotype; (137) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 162–173. Habitus: (162–165) Notopilo, cambageicola species group—(162) Notopilo cambageicola, paratype; (163) Notopilo interfabulatus, holotype; (164) Notopilo lawnhillensis, holotype; (165) Notopilo tanybasilaris, paratype; (166) Notopilo congruus (congruus species group), lectotype (of Opilo congruus); (167–168) Notopilo, reduncus species group—(167) Notopilo magnus, holotype; (168) Notopilo reduncus, paralectotype (of Opilo abdominalis Schenkling); (169–171) Notopilo, variipes species group—(169) Notopilo brevistriatus, paratype; (170) Notopilo gerstmeieri, paratype; (171) Notopilo variipes; (172–173) Notopilo, unplaced to species group—(172) Notopilo confusus, specimen from Western Australia; (173) Notopilo elstoni, holotype (scale bars: 1.0 mm).

FIGURES 181–185. (181) labels of the syntype specimen designated as the lectotype of Opilo congruus Newman (NHML); (182) labels of the syntype (holotype label invalid) specimen designated as the paralectotype of Opilo abdominalis Schenkling (IRSNB); (183) labels of the syntype specimen designated as the lectotype of Opilo variipes Chevrolat (MNHN); (184) labels of the syntype specimens designated as the lectotype of Opilo femoralis Westwood; (185) Westwood’s original Opilo femoralis syntype series (NHML) (a) O. femoralis lectotype, (b) smaller, non-conspecific, specimens representing a new species, Notopilo confusus sp. nov..