Notopilo beswickensis, Bartlett & Lambkin, 2022

Bartlett, Justin S. & Lambkin, Christine L., 2022, Australian Opilonini (Coleoptera: Cleridae: Clerinae) part I: A revised taxonomy for Australian Opilo Latreille including descriptions of new genera and species, Zootaxa 5220 (1), pp. 1-81 : 39

Notopilo beswickensis sp. nov.

ZooBank registration: urn:lsid:zoobank.org:act:D37A1778-FA33-40EB-9561-CBB61A918047

( Figs 43 , 81 , 116 , 156 ; Map 3)

HOLOTYPE ♁: Northern Territory: Austral. North.T. Beswick, I 1958, leg. H.Demarz // coll. R. Gerstmeier RGCM Munich ( QM, type reg. T258551) . PARATYPES (21): Northern Territory: same data as holotype (4, ZSM; 7, RGCM); Tindal, N.T. 14.31S 132.22E, 1-20 Dec. 1967, light trap, W.J.M. Vestjens (5, ANIC) . Queensland: Australia: n. Qld, 40 Mile Scrub N.P. via Mt Garnet , 10-i-25-ii-1986, Storey & Heiner // Malaise trap (1 ♁, 1, QDPC) ; Mareeba, N.Q., Jan 1950, C. Oke // COL-65682 (1, NMV) ; Mt Garnet Ra., N.Q., 1/58 GB // J.G. Brooks Bequest , 1976 (1, ANIC) . Western Australia: [unknown locality] W. Aust // F. E. Wilson Collection // Opilo near but not congruus // COL-65677 (1, NMV) .

Additional material: Northern Territory: Brunette Downs, NT, via Freewina , 12 February 1982, Light, R. Patterson, G 893 (1 ♀, ANIC) ; Tindal, N.T. 14.31S 132.22E, 1-20 Dec. 1967, light trap, W.J.M. Vestjens (1 ♀, ANIC) . Western Australia: WA: Argyle Diamond Mine , ca. 110 km SSW Kununurra 16°43′S 128°24′E 21 February 1984 A.C. Postle // Argyle Diamond Mines Collection donated 25 June 2001 (1, WAM) .

Diagnosis. Pronotum broadly rounded laterally, disc not heavily punctate, central sulcus small and shallow; elytra dark with white to pale yellow fasciate and apical maculations (humeral maculae absent in typical form), fasciae angled toward and meeting at suture, punctation with nodules, 8 th stria beginning at or within fascia, punctate to apex; legs entirely black, tarsi with three ventral tarsal pads.

Description. Habitus: Fig. 156 . Total length: 6–8.1 mm. Head: Cranium black, supra-antennal elevation and parts of clypeus often dark reddish-black, anteclypeus and labrum orange, palpi dark brown with paler terminal margins, antennae brown; frons broad (eyes separated by 1.65–1.8 eye widths), moderately punctate, rugulose above narrowest point, smoother below, slightly bulging above epistomal suture, clypeus with several punctures, anterior margin strongly concave, genae wrinkled; exterior margins of terminal palpomeres about 1.5–2 times (maxillae) and about 2.5 times (labium) the length of inside edges; antennae reaching base of pronotum; eyes and most of cranium vested with erect pale setae, frons with shorter medially-directed setae. Prothorax: Entirely black; pronotum 1.2–1.3 times longer than wide, moderately rotund in shape, anteriorly as broad as middle and about as wide or slightly narrower than head, sides round (not tuberculate), subapical depression deeply v-shaped, central sulcus weakly impressed; disc moderately to strongly punctate though smooth (glossy) in appearance, moderately distributed with short fine setae oriented in all directions and occasional long erect setae. Pterothorax: Ventrites dark brown, impunctate, moderately vested with short pale and occasional long setae; elytra blackish-brown with pale yellow to ivory-like markings (a large apical macula and a thick transverse fascia crossing the sutures and having the anterior and sometimes the posterior margins tapering inwards); elytra widest in apical half, length to width ratio 2.4–2.5:1; punctation with lateral nodules, nodules not visible within fascia, punctation large, deep and crowded in basal half, only slightly less so posterior of fascia, striae distinct until apical macula, punctation present but more confused within apical macula, 8 th stria beginning at or within fascia; interstriae with moderately long erect setae every 1 or 2 punctures and less frequently with longer erect setae, inner anterior rim of punctations with a minute semi-reclinate seta; epipleurae extending well into apical curve; hindwing with CuA 3+4 cross-vein complete (CuA 1 cross-vein not examined), MP 3+4 absent basad of CuA 1 crossvein. Legs: Entirely black to blackish-brown in most species (one female form Tindal with predominantly pale femora), tarsi dark brown, ventral tarsal pads paler; profemora barely swollen, other femora slender. Abdomen: Ventrites dark brown. Male genitalia: Tegmen ( Fig. 43 ) robust, sides subparallel, rounding off to parameroid lobes (apically tapered to a weakly digitiform process), dorsal sinus just over one-third tegmen length, weakly narrowing internally, ventral two-thirds as long, tegminal arms gradually meeting apodeme at broad base; median lobe as in Fig. 81 ; pygidium as in Fig. 116 .

Variation. The size and shape (degree of taper towards suture) of the elytral fascia is variable within specimens of the type series. Two non-type specimens, one from Brunette Downs, Northern Territory (ANIC), another from Argyle Diamond Mine , Western Australia (WAM), differ from those of the type-series in having a large humeral maculation on each elytron.

Etymology. The specific epithet beswickensis refers to Beswick in the Northern Territory where most of the type-series was collected.

Biology. Adults have been collected in December and January.

Distribution (Map 3). Specimens are known from northern parts of Queensland and the Northern Territory plus one unknown location in Western Australia.

Figures

FIGURES 27–47. Tegmina, dorsal: (27) Ancyropilus brigalowae, paratype; (28) Ancyropilus emmotti, holotype; (29) Ancyropilus exossuarius, paratype; (30) Ancyropilus monteithi, holotype; (31) Ancyropilus noonbahensis, holotype; (32) Ancyropilus packsaddlensis, holotype; (33) Ancyropilus simplex, paratype; (34) Ancyropilus tricolor, paratype; (35) Ancyropilus hornensis, paratype; (36) Culcipternotus mareebensis, paratype; (37) Infectostriatus absentis, holotype; (38) Infectostriatus differens, holotype; (39) Monilonotum bunyense, paratype; (40) Monilonotum doddi; (41) Monilonotum pascoei; (42) Monilonotum sundholmi, paratype; (43–47) Notopilo, beswickensis species group—(43) Notopilo beswickensis, holotype; (44) Notopilo calicis, paratype; (45) Notopilo tompricensis, holotype; (46) Notopilo xanthoimprocerus, holotype; (47) Notopilo xanthoprolatus, holotype (scale bars: 0.5 mm).

FIGURES 66–101. Median lobes: (66) Ancyropilus brigalowae, paratype; (67) Ancyropilus emmotti, holotype; (68) Ancyropilus exossuarius, paratype; (69) Ancyropilus monteithi, holotype; (70) Ancyropilus noonbahensis, holotype; (71) Ancyropilus packsaddlensis, holotype; (72) Ancyropilus simplex, paratype; (73) Ancyropilus tricolor, holotype; (74) Ancyropilus hornensis, paratype; (75) Culcipternotus mareebensis, paratype; (76) Infectostriatus absentis, holotype; (77) Infectostriatus differens, holotype; (78) Monilonotum bunyense, paratype; (79) Monilonotum pascoei; (80) Monilonotum sundholmi, paratype; (81–85) Notopilo, beswickensis species group—(81) Notopilo beswickensis, holotype; (82) Notopilo calicis, paratype; (83) Notopilo tompricensis, holotype; (84) Notopilo xanthoimprocerus, paratype; (85) Notopilo xanthoprolatus, holotype; (86–88) Notopilo, cambageicola species group—(86) Notopilo cambageicola, paratype; (87) Notopilo interfabulatus, paratype; (88) Notopilo tanybasilaris, paratype; (89) Notopilo congruus (congruus species group); (90) Notopilo reduncus (reduncus species group); (91–93) Notopilo, variipes species group—(91) Notopilo brevistriatus, paratype; (92) Notopilo gerstmeieri, holotype; (93) Notopilo variipes; (94–96) Notopilo, unplaced to species group—(94) Notopilo elstoni, holotype; (95) Notopilo katherinensis, holotype; (96) Notopilo obesus, paratype; (97) Platynotum bulli, paratype; (98) Platynotum culgoense, holotype; (99) Platynotum femorale; (100) Platynotum foveosetosa, holotype; (101) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 102–137. Male pygidiae: (102) Ancyropilus brigalowae, paratype; (103) Ancyropilus emmotti, holotype; (104) Ancyropilus exossuarius, paratype; (105) Ancyropilus monteithi, holotype; (106) Ancyropilus noonbahensis, holotype; (107) Ancyropilus packsaddlensis, holotype; (108) Ancyropilus simplex, paratype; (109) Ancyropilus tricolor, paratype; (110) Ancyropilus hornensis, paratype; (111) Culcipternotus mareebensis, paratype; (112) Infectostriatus absentis, holotype; (113) Monilonotum doddi; (114) Monilonotum pascoei; (115) Monilonotum sundholmi, paratype; (116–120) Notopilo, beswickensis species group—(116) Notopilo beswickensis, holotype; (117) Notopilo calicis, paratype; (118) Notopilo tompricensis, holotype; (119) Notopilo xanthoimprocerus, paratype; (120) Notopilo xanthoprolatus, holotype; (121–124) Notopilo, cambageicola species group—(121) Notopilo cambageicola, paratype; (122) Notopilo interfabulatus, paratype; (123) Notopilo lawnhillensis, paratype; (124) Notopilo tanybasilaris, holotype; (125) Notopilo congruus (congruus species group); (126–128) Notopilo, variipes species group—(126) Notopilo brevistriatus, paratype; (127) Notopilo gerstmeieri, holotype; (128) Notopilo variipes; (129–132) Notopilo, unplaced to species group—(129) Notopilo confusus, holotype; (130) Notopilo elstoni, holotype; (131) Notopilo katherinensis, holotype; (132) Notopilo obesus, paratype; (133) Platynotum bulli, paratype; (134) Platynotum culgoense, holotype; (135) Platynotum femorale; (136) Platynotum foveosetosa, holotype; (137) Platynotum gracile, holotype (scale bars: 0.5 mm).

FIGURES 150–161. Habitus: (150) Infectostriatus differens, holotype; (151) Monilonotum bunyense, paratype; (152) Monilonotum doddi; (153) Monilonotum pascoei; (154) Monilonotum sundholmi, paratype; (155) Monilonotum rufiventre; (156–161) Notopilo, beswickensis species group—(156) Notopilo beswickensis, holotype; (157) Notopilo calicis, holotype; (158) Notopilo eremosus, holotype; (159) Notopilo tompricensis, holotype; (160) Notopilo xanthoimprocerus, holotype; (161) Notopilo xanthoprolatus, holotype (scale bars: 1.0 mm).