Anaspis bertrami, Levey, 2020

Levey, Brian, 2020, A review of the species of Anaspis (s. str.) similar to A. nigripes Brisout and A. apfelbecki Schilsky, with the description of three new species (Coleoptera: Scraptiidae), Zootaxa 4778 (3), pp. 509-520 : 511-513

Anaspis bertrami sp. n.

( Figs 5 , 11 , 20 , 28 )

Type specimens. Holotype ♂ ( NMGW). N. Lebanon: Horsh Ehden Nature Reserve , 34.19′ N 36.00 ′E, 1,300–1,600m, 29.v.–4.vi.2001. B.Levey. NMWZ2000 081 ./ Holotype Anaspis bertrami sp.n. B.Levey2008. Paratypes: 2♀ ( NMGW, BMNH), same data as Holotype, but one with extra label:Lower part of reserve with Pinus spp. Quercus spp.and broom .

Diagnosis. General diagnosis of holotype (male): Length (fore margin of pronotum to apex of elytra in mid line) 3.6 mm. Head above antennal insertions, pronotum, scutellum, elytra and underside black; frons opposite antennal insertions, labrum, basal three segments of maxillary palpi, basal five segments of antennae, coxae, trochanters and basal three-quarters of femora of fore legs, basal half of mid femora, and tibial spurs of all legs yellow; apical six segments of antennae black; rest of fore legs yellowish brown, remainder of mid and hind legs blackish brown. Entire body densely covered with short grey pubescence.

Head. Antennae: first antennomere almost twice as long as wide, nearly rectangular; second antennomere cylindrical, slightly longer than first antennomere; third and fourth antennomeres about equal length, cylindrical, slightly widened at apex, about 1.5× as long as the second; fifth and sixth antennomeres about equal length, cylindrical, slightly shorter and broader than fourth; seventh about equal to sixth, slightly triangular; eighth to tenth antennomeres becoming progressively shorter and broader, nearly triangular; eleventh ovate, about 1.3× as long as 10. Apical segment of maxillary palp securiform.

Pronotum: about 1.3× as wide at base as long in mid line; hind angles obtuse, rounded; lateral edge straight when viewed from side. Scutellum: triangular about as wide as long. Elytra: 3.4× as long as pronotum; 2.8× as long as wide over the humeri. Abdomen ( Fig. 20 ): first and second ventrites convex, but not keeled; third ventrite triangular in cross section, with a sharp median keel, with a small projecting tuft of posteriorly directed setae in mid line and a pair of long slightly curved, distally twisted, appendages, widely separated at the base; fourth ventrite weakly convex, the posterior margin narrowly, weakly emarginated, almost truncate medially; fifth ventrite deeply emarginated at centre. Legs: fore leg ( Fig. 5 ) femora very broad and strongly flattened anterior-posterior and slightly excavated on the posterior surface; tibia of the same length as femora, slightly flattened anterior-posterior, slightly widening distally with a dense distal fringe of dark brown spines. Tibial spurs present, yellow coloured, the longest slightly longer than the second tarsal segment; tarsal segment 1 small, about as wide as long, slightly widening distally; 2 about 1.5× as long as 1, as wide at base as 1 at apex, widening distally, about as wide as long, with a ventral pad extending distally beyond the base of the segment 3; 3 about as long as 2, approximately heart shaped with a deep medial excision distally, about 1.25× as wide at apex as long, with a long semicircular ventral pad extending distally beneath the entire length of segment 4; 4 very small oblong, slightly longer than wide; 5 elongate, club shaped about as long as 3 and 4 together, claws small curved, widened at base. Mid leg with femora moderately anterior-posterior flattened; tibia about as long as femora, regularly widening from base to apex, the ventral margin straight; with a dense distal fringe of yellow-brown spines and a pair of long yellow spurs, the longer about twothirds length of tarsal segment 1, the shorter about half length of segment 1; tarsal segment 1 long, cylindrical, 0.55 length of tibia; 2 shorter, cylindrical, 0.50 length of 1; 3 shorter, 0.50 length of 2, approximately heart shaped with a deep medial excision, with a long semicircular ventral pad; remaining segments lost. Hind legs missing. Genitalia ( Fig. 11 ): parameres hooked at apex; basal piece 1.3× as long as parameres.

General diagnosis of ♀ ( Fig. 28 ): Length 3.80–3.90 mm. Colour as in male but pronotum orange-yellow, and frons from below antennal insertions and labrum yellow. Legs coloured as male in one female, other female with legs entirely blackish brown except for the yellow tibial spurs. Head: same as male but segments 7–10 of antennae slightly shorter and broader. Pronotum: same as male. Scutellum: same as male. Elytra: 3.7× as long as pronotum; 2.8–2.9× as long as wide over the humeri. Abdomen: ventrites 1–4 convex not keeled; ventrite 5 approximately triangular with a rounded apex. Legs: fore leg with femora strongly flattened anterior-posterior; tibia about twothirds length of femora, slightly flattened anterior-posterior, slightly widening distally with a dense distal fringe of yellow-brown spines and two yellow spurs the longest about half length of tarsal segment 1; tarsal segment 1 elongate, slightly widening distally, about 3.0× as long as wide; 2 about two-thirds length of 1, and about as broad as 1; 3 about as long as 2, approximately heart shaped with a deep medial excision distally, about 1.25× as wide at apex as long, with a long semicircular ventral pad extending distally beneath the entire length of segment 4; 4 very small oblong, slightly longer than wide; 5 elongate, club shaped about as long as 3 and 4 together, claws small curved, widened at base. Mid leg as in male but tibial spurs and tarsi slightly shorter relative to the length of the tibia. Hind leg with femora strongly flattened anterior-posterior; tibia about half length of femora, strongly widened from base to apex, about 3.0× as long as wide at apex, with a dense distal fringe of short brown spines and two yellow spurs, the longer about two-thirds the length and the shorter about half the length of tarsal segment 1; tarsal segment 1 moderately widening at apex, about four× as long as wide at apex, about 0.70 length of tibia; 2 slightly widening at apex, about 0.70 length of 1; 3 very slightly widening at apex, about 0.70 length of 2; 4 cylindrical, about as long as 3, claws small, slightly curved, slightly widened at base.

Comparative diagnosis. This species is closely related to A. curva sp. n. differing mainly in the form of the appendages of ventrite 3 of the male, which are less strongly curved, twisted distally, with the grooved face facing ventrally ( Fig. 20 ) (appendages, more strongly curved, not twisted distally, with the grooved face inward facing in A. curva sp. n. ( Fig.19 )).

Etymology. I name this species in memory of my father Bertram Levey in recognition of his support, which gave me the opportunity to become a biologist.

Figures

FIGURES 1–9. Fore legs of male: 1. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 2. A. nigripes Brisout de Barneville, France, Côte-d’Or, Pouilly –en-Auxois (NMGW); 3. A. cooteri sp. n., holotype (NMGW); 4. A. curva sp. n., holotype (NMGW); 5. A. bertrami sp. n., holotype (NMGW); 6. A. graeca Schilsky, syntype (BMNH); 7. A. apfelbecki Schilsky, paralectotype (ZMHB); 8. A. apfelbecki Schilsky, paralectotype (ZMHB); 9. A. marseuli Csiki, Japan, Tochigi, Tsurutanuma, Utsunomiya City (TPMJ).

FIGURES 10–17. Male genitalia: 10. A. curva sp. n., holotype (NMGW); 11. A. bertrami sp. n., holotype (NMGW); 12. A. apfelbecki Schilsky, paralectotype (ZMHB); 13. A. apfelbecki Schilsky, paralectotype (ZMHB); 14. A. cooteri sp. n. holotype (NMGW); 15. A. nigripes Brisout de Barneville, France, Côte-d’Or, Pouilly-en-Auxois (NMGW); 16. A. koenigi Schilsky, paralectotype (ZMHB); 17. A. marseuli Csiki, Japan, Tochigi, Yunishikawa, Kuriyama Vill. (TPMJ).

FIGURES 18–27. Apical abdominal ventrites of male: 18. A. apfelbecki Schilsky, paralectotype (ZMHB); 19. A. curva sp. n., holotype; 20. A. bertrami sp. n., holotype (NMGW); 21. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 22. A. nigripes Brisout de Barneville, Spain: La Rioja, Nieva de Cameros, La Dehesa (NMGW); 23. A. koenigi Schilsky, paralectotype (ZMHB); 24. A. marseuli Csiki, Japan, Tochigi City (TPMJ); 25. A. marseuli Csiki, Japan, Utsunomiya City (TPMJ); 26. A. cooteri sp. n., paratype (NMGW); 27. A. graeca Schilsky, syntype (BMNH).

FIGURES 28–36. Dorsal habitus: 28. Anaspis bertrami sp. n. female paratype (NMGW); 29. A. curva sp. n., male holotype (NMGW); 30. A. emarginata Schilsky, N. China, Heilunkiang, Erlungshan (BMNH); 31. A. cooteri sp. n., male paratype (NMGW); 32. A. marseuli Csiki, Japan, Hyogo, Haruki, Onsen Town (TPMJ); 33. A. nigripes Brisout de Barneville, male, France, Côte-d’Or, Pouilly-en-Auxois (NMGW); 34. A. apfelbecki Schilsky, paralectotype (ZMHB); 35. A. koenigi Schilsky, paralectotype (ZMHB); 36. A. graeca Schilsky, syntype (BMNH).