Tonkinospira Jochum

Páll-Gergely, Barna, Jochum, Adrienne & Asami, Takahiro, 2017, Three New Species And A New Genus Of Hypselostomatidae (Gastropoda: Pulmonata) From Cong Troi Cave, Northern Vietnam, Acta Zoologica Academiae Scientiarum Hungaricae 63 (3), pp. 327-341 : 337-340

Tonkinospira Jochum , SlapniketPáll- Gergely, 2014

Systenostoma Bavay & Dautzenberg, 1908: JournaldeConchyliologie, 56: 243. not Systenostoma Marsson, 1887 [Bryozoa]. Establishedasasubgenusof Helix .

Systenostoma — Bavay & Dautzenberg, 1908, JournaldeConchyliologie, 57: 196. (diagnosis).

Tonkinospira Jochum , SlapniketPáll- Gergely, 2014: In: JOCHUMetal., ZooKeys 410: 31.

Typespecies: Helix (Systenostoma) pauperrima BavayetDautzenberg, 1908

bysubsequentdesignation (PILSbRY 1917).

Content: defixa (BavayetDautzenberg, 1912) , depressa ( Jaeckel, 1950) , pauperrima (BavayetDautzenberg, 1908) , pulverea (BavayetDautzenberg, 1908) , tomasini Páll- GergelyetJochum, sp. n.

Distribution: Tonkinospira speciesareknownfromNorthernVietnamso far ( PÁLL-GERGELY et al. 2015 and this study).

Tonkinospiratomasini Páll- GergelyetJochumsp. n.

( Figs 1A, 5, 6, 7A–B)

Typematerial: C03-N39 (localitycode), NorthernVietnam, HaGiangProvince, Quan BaDistrict, TamSon, CongTroiCave, 980 m, 23°02’44.7”N, 104°59’23.9”E, WGS84, atthe entrance, goursofdrippingwater, leg. G. TOMASIN, 28.01.2010., Specimen1 (= holotype, MZUFGC/49399); Samedata, Specimen2 (figuredparatype), Specimen3 (figuredpara- type), additional 4 paratypes, MZUFGC/49400; Samedata, HNHM 99731 / 1 paratype ; Samedata, PGB / 1 paratype ; Samedata, 2 juvenileshells (notparatypes), MZUFGC/49401; C04-N74 (localitycode), NorthernVietnam, HaGiangProvince, QuanBaDistrict, Tam Son, CongTroiCave, 980 m, 23°02’44.7”N, 104°59’23.9”E, WGS84, internalcanyon, gours ofdrippingwater, leg. G. TOMASIN, 29.01.2010., MZUFGC/49407/ 1 paratype.

Diagnosis: Smallconicalshellwithconvexwhorls, lessthan 1.95 mm

broad; apertureedentate.

Description: Shellconvexlyconicwithobtuseapex, greyishwhite; nuclearwhorlbe- ginswithapatternofgranulosewrinkles, mergingintoapatternofdense, shallowpits seeTable 1.

Fig. 6. Variation of shell shape in Tonkinospira tomasini sp. n. A = n39, Specimen2, B = n39, Specimen1 (holotype); C = n39, Specimen3

mellar, 5–6 = palatal

withoutspiralthreads, granulartexture; suturedeep; protoconchconsistsofapproximate- ly 1.5 whorls; teleoconchretiformwithfaintspiralthreads; protoconchandteleoconchcon- sistsof 5 evenlyincreasingconvexwhorls; aperturequadrate- reniform, adnate; leftouter margin thickly angular with decreasing thickness descending towards the basal margin andascendingtowardstheupperrightportionoftheperistome; basal- mostsectionof peristomefoldsbackslightly; distalendofthebodywhorldoesnotproceedpastpenul- timatewhorlbutratheralignsevenwiththeventralconvexityofthepenultimatewhorl inprofileview; distalendofbodywhorldeflectstotherightoftheumbilicus; umbilicus narrow and deep.

Measurements (in mm): SH = 2.23–2.98; SW = 1.7–1.93; AH = 0.79–0.91; AW = 0.82– 0.94 (n = 8) ( Table 3).

Differentialdiagnosis: Themostsimilar Tonkinospira speciesis T. pauperrima , which has a wider shell, a lower spire and shows a more pronounced spiralstriationthan T. tomasini sp. n.

Fig. 8. Distributionof Angustopila species. 1 = typelocalityof A. huoyani , 2 = otherlocality of A. huoyani (see PÁLL-GERGELY et al. 2015), 3 = type locality of A. dominikae , A. subelevata and A. szekeresi , 4 = type locality of A. stochi sp. n. (Cong Troi Cave) and other locality of A. subelevata , 5 = typelocalityof A. fabella , 6 = typelocalityof A. tamlod , 7 = typelocalityof A.

elevata , 8 = type locality of A. concava

Etymology: ThenewspeciesisnamedafterG. Tomasin, whocollectedthematerial

fromCongTroiCave.

Distribution: Tonkinospiratomasini sp. n. isknownfromsedimentsam-

plesfromtheCongTroiCaveinNorthernVietnam.