Eulichas baeri ( Fairmaire, 1898 )

Hájek, Jiří, 2009, Revision of the genus Eulichas Jacobson, 1913 (Coleoptera: Eulichadidae) II. E. dudgeoni species group, Zootaxa 2192, pp. 1-44 : 24-26

Eulichas baeri ( Fairmaire, 1898)

( Figs. 8–9 , 32 , 52 , 72 )

Lichas baeri Fairmaire, 1898: 388 (original description, Luzon).

Eulichas baeri ( Fairmaire, 1898) : Pic 1914: 11 (catalogue); Jäch 1995: 363 (catalogue). Eulichas baeri var. innotatus Pic, 1924: 31 (original description, Luzon); Jäch 1995: 363 (catalogue). syn. nov.

Type locality. “Luzon” [ Philippines] ( E. baeri ), “Luzon” [ Philippines] ( E. baeri innotatus ).

Type material. Lichas baeri : not studied, no syntype was found in Fairmaire’s collection ( MNHN). Eulichas baeri var. innotatus : Lectotype 3 ( MNHN), by present designation: “Mt. Makiling / Luzon, Baker [printed] // 21160 [handwritten] // L. Baeri / Frm. var. [handwritten] // Muséum Paris / Coll. M. Pic [printed] / / TYPE [red label, printed] // v. innotatus / mihi [handwritten] // LECTOTYPUS / EULICHAS / baeri / var. innotatus Pic, 1924 / Jiří Hájek des. 2002 [red label, printed] // EULICHAS (s. str.) / baeri ( Fairmaire, 1898) / Jiří Hájek det. 2002 [printed]”. Number of syntypes unknown. I designate a lectotype to fix the identity of this species, as available taxonomic works do not allow an unambiguous identification of specimens.

Additional material studied. 155 specimens — BRUNEI: 13, Bukit Teraja, Labi [ca. 04°24’N 114°27’E], Dipterocarp. forest, light trap, 60 m, 2.viii.1979, S.L. Sutton leg. ( BMNH); 233 1Ƥ, Temburong, Kuala Belalong F.S.C., mixed Dipterocarp forest, 60–300 m, 16.–20.iv.1993, E. Heiss leg. ( NHMW). INDONESIA: 1Ƥ, Kalimantan Timur, Boven, Mahakkam, 1894, Dr. Nieuwenhuis exped. ( RMNH); 13, Kalimantan Tengah, Busang / Rekut confluence, 0°03’S 113°59’E, light trap, M.J.D. Brendell & H. Mendel leg. ( BMNH); 13, Kalimantan Timur, Sanga Sanga, H.D. Jensen leg. ( BMNH). MALAYSIA: SABAH: 433 7ƤƤ, Kinabalu ( HNHM, IRSNB, MNHN, SMTD); 13 1Ƥ, Mount Kinabalu ( NHMB); 233 4ƤƤ, same label data, but v.–viii.1903, J. Waterstradt ( MNHN); 1Ƥ, Kinabalu NP, Head Q., 06°00’N, 116°32’E, 1600 m, nearly undisturbed montane evergreen rain forest, 15.v.1987, J. Huisman leg. ( RMNH); 2ƤƤ, Kinabalu Nat. Park, Sayap, 1000 m, 25.–29.xi.1996, W. Schawaller leg. ( NMPC, SMNS); 3ƤƤ, same label data, but 6°10’N 116°34’E, primary forest edge, 950 m, at light, 8.iii.2001, J.P. Duffels & M.A. Schouten leg. ( ZMAN); 1Ƥ, Crocker Range, 16 miles NW of Keningau, 4.iv.1982, S. Nagai leg. ( EUMC); 433 8ƤƤ, Crocker Range Mts., 50 km E Kota Kinabalu, Gunung Emas, 16.–27.iv.1993, I. Jeniš leg. ( NHMW); 13 same label data, but 500–1900 m, 5.–21.v.1995, J. Stolarczyk leg. ( NMPC); 13 2ƤƤ, Banjaran Crocker Mts., 16 km SW Gunung Alab, 790–850 m, 4.–9.v.1996, M. Štrba & R. Hergovits leg. ( NHMW); 1Ƥ, 105 km S Beaufort, Long Pa Sia area, airstrip Long Pasia, 04°24’N, 115°43’E, 1000 m, at light, 15.iv.1987, v. Tol & Huisman leg. ( RMNH); 13, 65 km W Lahad Datu, Sg. Purut camp, 04°57’N, 117°45’E, 250 m, at light, 26.x.1987, J. Huisman & R. de Jong leg. ( RMNH); 13, 15 km W of Lahad Datu, Danum valley, light trap, 7.xii.1989, M.J. & J.P. Duffels leg. ( ZMAN); 13, 70 km W Lahad Datu, Danum valley, Next to bridge Nature Trail, 150 m, rainforest along narrow stream, at light, M.J. & J.P. Duffels leg. ( NMPC); 1Ƥ, same label data, but 13.xii.1989 ( ZMAN); Poling, near Ranau, 29.iv.1980, M. & A. Sakai leg. ( EUMC); 13, Ranau Distr., Poring Hot Spring, Poring Lodge, 06°02’35’’N, 116°42’19’’E, ca. 650 m, 14.–16.v.2002, T. Kothe leg. ( ZSMC); 1Ƥ, West Coast zone, 12 km NNE Ranau, Poring Hot Springs, 06°04’N, 116°42’E, 550 m, at light, 9.–10.xi.1987, J. Huisman & R. de Jong leg. ( RMNH); 2 33, Poring Hot Spring, 450–600 m, 9.–11.iii.2007, W. Schawaller leg. ( SMNS); 13, 80 km E of Ranau, Telpid, 800 m, xii.1978, S. Nagai leg. ( EUMC); 13 1Ƥ, Tawau [ca. 04°15’N 117°52’E], 20.iii.2006 ( NMPC). SARAWAK: 2ƤƤ, 4th Division, Gunung Mulu National Park, at light, iii. 1978, J.D. Holloway et al. leg. ( BMNH); 333 1Ƥ, same label data, but 150–200 m, v.–viii.1978, P.M. Hammond & J.E. Marshall leg. ( BMNH); 13, Rumah Kabau anak muggot, Ng Sebong Baleh, 25 km E Kapit, iii.1994, J. Kodada leg. ( NHMW); 13, Kapit distr., Rumah Ugap vill., Sut. riv., 3.–9.iii.1994, S. Bílý leg. ( NMPC); 13, Kapit distr., Sebong, Baleh river, 9.–21.iii.1994, J. Horák leg. ( NHMW). PHILIPPINES: LEYTE: 13 ( NMPC); 2ƤƤ, 22.viii.2003, J.L. Boundant leg. ( MCSN); 13, Cabalian [= San Juan, ca. 10°16’N 125°09’E], coll. W. Schultze ( SMTD); 13, Mt. Balocaue, 700 m, local collector leg. ( NMPC). LUZON: 13, Mt. Isarog ( SMTD); 1Ƥ, Pangasinan [most probably = Pangasanan, ca. 14°58’N 120°42’E], coll. Benesh ( SMTS); 2ƤƤ, Sorsogon [ca. 12°58’N 124°00’E] ( NHMB); 1Ƥ, Upper Bintacan, Isabela, 30.iv.1983, S. Osada leg. ( EUMC); 13, Nueva Viscaya, Sta Fe, datton Pass, 900 m, at light, 3.–8.vi.1985, R.A. Müller leg. ( RMNH); 233 1Ƥ, Nueva Vizcaya, vi.2003, I.A. Lumawig leg. ( MCSN, NMPC). MINDANAO: 1Ƥ, coll. W. Schultze ( SMTD); 13, Davao [ca. 07°04’N 125°36’E], Dr. Platen ( MNHN); 13, Davao, iii.1968 ( EUMC); 1Ƥ, Davao, 5.xi.1991 ( MCSN); 1Ƥ, Davao prov., Baracatan, Mt. Talomo, 1100 m, 3.–6.viii.1985, M. Owada leg. ( EUMC); 1Ƥ, Upper Barakatan, Apo Range, 1100 m, 17.viii.1985, M. Tomokuni leg. ( EUMC); 1Ƥ, Baracatan, Eagle Centre, N slopes of Mt. Apo, 1100 m, 5.viii.1985, M. Sakai leg. ( EUMC); 1Ƥ, Agko, Mt. Apo [ca. 07°01’N 125°16’E], 1000 m, 7.x.1978, S. Nagai leg. ( EUMC); 2ƤƤ, same label data, but 9.iii.1979 ( EUMC); 13, Mt. Apo, 8.–10.i.1981, Natives leg. ( EUMC); 233, Mt. Apo, iv.2005, I.A. Lumawig leg. ( MCSN); 13, Lindaban [ca. 8°17’N 124°48’E], Bukindon, W. Schultze leg. ( SMTD); 13, Manila, coll. Schwarz ( DEIC); 1Ƥ, Surigao prov., W. Boettiker leg. ( SMTD); 13, Surigao [ca. 9°47’N 125°28’E], Baker leg. ( SMTD); 13, Surigao, W. Schultze leg. ( SMTD); 233 2ƤƤ, Surigao, 15.v.1915 ( NMPC, ZMHB); 1Ƥ, Tudaya-Sibulan, 30.vii.1970, T. Okadome leg. ( EUMC); 1Ƥ, Gasy, Maitum, S Cotabato, 700 m, 12.viii.1985, M. Sakai leg. ( EUMC); 2ƤƤ, South Cotabato, Parker Mts., Salacale, 600–900 m, at light, 2.–12.iv.1985, R.A.Müller leg. ( RMNH); 13, South Cotabato, Koronadal Barrio 8, 100– 300 m, 12.–14.vii.1986, R.A. Müller leg. ( RMNH); 13 1Ƥ, Bagong Silang, 30 km NW Maramag, 1700 m, 13.–17.v.1996, BOLM leg. ( NMPC, SMTD); 733 3ƤƤ, Davao Oriental, Boston, Agtuuganon Mt., Camp 55, 1020 m, 29.-v.-7.vi.1997, Müller, Buenafe & Gorost. leg. ( NMPC, RMNH); 13, Surigao del Sur Lianga, 8 km W Diatagon, 08°42’N 126°05’E, 200 m, 3.–7.vii.2005, J.H. Lourens leg. ( BMNH); 13 2ƤƤ, Agusan Sur, 10 km SE Trento Sta Maria, 08°01.615’N, 126°12.322’E, 185 m, 27.–28.iv.2008, J.H. Lourens leg. ( IRSNB); 13, Davao Oriental, Aliwagwag Primary forest, 07°43.667’N, 126°17.304’E, 90 m, 30.iv.–1.v.2008, J.H. Lourens leg. ( IRSNB). MINDORO: 333, Abra de Ilog [ca. 13°27’N 120°43’E], W. Schultze leg. ( NMPC, SMTD). NEGROS: 13, vi.1991 ( MCSN); 13, v.1996 ( NMPC). PALAWAN: 233 1Ƥ, Cleopatra Needle Nat. Park, Tanabag river valley, 300 m, 20.–22.xii.1990, BOLM leg. ( NHMB, NMPC). SAMAR: 13, Baker leg. ( MNHN); SIBUYAN: 13 1Ƥ, Romblon, vii.1985 ( EUMC); 1233 2ƤƤ, Romblon prov., Magdiwang, Pawala river, Tampayan Camp, Ga-ong area, 70–400 m, at light, 19.iii.–7.iv.1987, R.A. Müller leg. ( NMPC, RMNH).

Description. Habitus elongate, fusiform. Body colouring from brownish-red to brown. Pale part of setation consists of recumbent yellowish or greyish setae covering uniformely body surface, or forming more or less distinct ocellations on elytra ( Figs. 8–9 ).

Measurements. Males: 13–24 mm; females: 20–26 mm.

Head punctation consists of sparse, irregularly distributed moderately large setigerous punctures. Antenna slender, last antennomere rather oval, 2.57–3.58 times as long as wide ( Fig. 32 ), its ventral side smooth.

Pronotum trapezoidal, ca. 1.71–1.93 times as wide as long. Sides of rather variable shape. In some specimens skewed and straight in anterior half, with indistinct angle near the middle, and hind angles distinctly prominent ( Fig. 52 ). On the other hand almost regularly rounded in some specimens. The disc with two indistinct rounded depressions. Punctation consists of sparse moderately coarse setigerous punctures on the disc, which become slightly coarser and denser laterally.

Elytra with numerous longitudinal rows of moderately large punctures, and very fine interstitial setigerous punctures.

Ventral part with fine punctures, which are sparse medially and become larger and denser laterally. Last abdominal ventrite laterally almost regularly rounded to apex.

Male. Aedeagus with phallobase of the same length as parameres. Parameres simple, relatively broad, subparallel. Parameral subbasal hook is missing, but the subapical hook is developed. Median lobe narrowly lanceolate ( Fig. 72 ).

Female. Similar to male in habitus, but larger. Antenna shorter, and more slender.

Remarks. So far the only Eulichas species known from the Philippines, thus I was able to identify it, although no syntype was found/recognised in Fairmaire’s collection (MNHN). A variable species. The variety innotatus represents only indistinctly ocellated specimens. It is without any doubt conspecific with typically coloured specimens, and is therefore junior subjective synonym of E. baeri . However, populations from different Philippine islands, as well as from Borneo, differ in body size, shape of last antennomere and pronotum. In particular, specimens from Borneo have usually more rounded sides of pronotum and more distinct ocellations on elytra, and may represent a separate subspecies. However, as the same characters can also be observed in some specimens from different Philippine Islands, I refrain from its formal description.

Collection circumstances. Collected at light.

Distribution. Known from the Greater Sunda Island of Borneo – Brunei, Indonesia (Kalimantan), Malaysia (Sabah, Sarawak), and the Philippines (Leyte, Luzon, Mindanao, Mindoro, Negros, Palawan, Samar).

Figures

FIGURES 1 – 12. Habitus of Eulichas. 1 — E. alesbezdeki sp. nov. (holotype); 2 — E. bertiae (paratype, Suisapa); 3 — E. dudgeoni (paratype, Hong Kong); 4 — E. jendeki (Sa Pa); 5 — E. siamensis sp. nov. (paratype, Ban Bo Klua); 6 — E. similis sp. nov. (paratype, Soppong); 7 — E. uniformis (Dolakha); 8 — E. baeri (Luzon); 9 — E. baeri (Crocker Range); 10 — E. gigantea (holotype); 11 — E. incisicollis ♂ (Tanah Rata); 12 — E. incisicollis Ƥ (holotype).

FIGURES 25 – 44. Male antennomeres IX – XI of Eulichas. 25 — E. alesbezdeki sp. nov.; 26 — E. bertiae; 27 — E. dudgeoni; 28 — E. jendeki; 29 — E. siamensis sp. nov.; 30 — E. similis sp. nov.; 31 — E. uniformis; 32 — E. baeri; 33 — E. incisicollis; 34 — E. oborili sp. nov.; 35 — E. robusta sp. nov.; 36 — E. sausai sp. nov.; 37 — E. serricornis sp. nov.; 38 — E. sikkimensis; 39 — E. sundaensis sp. nov.; 40 — E. wewalkai sp. nov.; 41 — E. fasciolata; 42 — E. jakli sp. nov.; 43 — E. subocellata; 44 — E. villosa sp. nov. Scale bar 1 mm.

FIGURES 45 – 54. Pronotum of Eulichas. 45 — E. alesbezdeki sp. nov.; 46 — E. bertiae; 47 — E. dudgeoni; 48 — E. jendeki; 49 — E. siamensis sp. nov.; 50 — E. similis sp. nov.; 51 — E. uniformis; 52 — E. baeri (Luzon); 53 — E. incisicollis; 54 — E. oborili sp. nov. Not in Scale.

FIGURES 72 – 77. Aedeagus of Eulichas in dorsal view. 72 — E. baeri; 73 — E. incisicollis; 74 — E. oborili sp. nov.; 75 — E. robusta sp. nov.; 76 — E. sausai sp. nov.; 77 — E. serricornis sp. nov. Scale bar 1 mm.