Geodromicus spadiceus Shavrin, 2019

Shavrin, Alexey V., 2019, The crassipalpis species group of the genus Geodromicus Redtenbacher, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini), Zootaxa 4686 (4), pp. 571-580 : 576-577

Geodromicus spadiceus Shavrin , sp.n.

( Figs. 3 , 6–7 , 11 )

Type material examined: Holotype ♂: ‘ Nepal Manaslu Mts. | N-slope Samdo | Sanam Khola | 3900m 1.VI.2006 | leg. J. Schmidt’ <rectangular label, printed >, ‘ HOLOTYPE | Geodromicus | spadiceus sp.n. | Shavrin A.V. des. 2019’ <red rectangulat label, printed> ( NME) .

Paratypes (7 specimens): 3 ♂♂ [two specimens dissected], 4 ♀♀ [two specimens without right antennomeres 9–11 and 7–11 respectively]: same data as the holotype, with additional red rectangular printed label : ‘ PARATYPE | Geodromicus | spadiceus sp.n. | Shavrin A.V. des. 2019’ (1 ♂, 1 ♀: CS; 2 ♂♂, 3 ♀♀: NME) .

Description. Measurements (n=8): HW: 1.13–1.26; HL: 0.76–0.84; OL: 0.27–0.28; TL: 0.08; AL (holotype): 3.87; PL: 0.94–0.97; PWmax: 1.44–1.66; PWmin: 1.32–1.47; ESL: 2.09–2.27; EW: 2.31–2.72; MTbL (holotype): 1.65; MTrL (holotype): 0.77 (MTrL 1–4: 0.45; MTrL 5: 0.32); AW: 2.13–2.68; AedL: 1.35; BL: 5.87–7.30 (holotype: 6.90).

Head, antennae, mouthparts, pronotum, femora and abdomen brown; elytra reddish-brown, usually with darker prescutellar area; apex of apical portion of maxillary palpi, sometimes apical antennomeres and tibia yellow-brown; tarsi yellow. Median portion of head with or without indistinct, irregular transverse meshes, denser between ocelli; infraorbital ridges with indistinct fine to distinct isodiametric microsculpture; pronotum with fine transverse microreticulation, more distinct in anterio-lateral and markedly sparser in posterio-median portions, sometimes with small impunctate area in posterio-median third. Habitus as in Fig. 3 .

Head 1.4–1.5 times as wide as long, with distinctly convex infraorbital ridges; anteromedian depression deep and wide, gradually narrowed basad and extending toward very wide and moderately deep interocellar depression. Ocelli very small, sometimes indistinct; distance between ocelli markedly longer than distance between ocellus and posterior margin of eye. Punctation irregular, moderately coarse, smooth, markedly denser on infraorbital ridges. Apical maxillary palpomere 1.7 times as long as preapical segment. Antennae very long, almost reaching apical margin of elytra when reclined; lengths × widths of antennomeres (holotype): 1: 0.37 × 0.15; 2: 0.27 × 0.10; 3: 0.37 × 0.12; 4–5: 0.32 × 0.12; 6–9: 0.35 × 0.12; 10: 0.32 × 0.12; 11: 0.50 × 0.12.

Pronotum 1.5–1.7 times as wide as long, 1.2–1.3 times as wide as head, widest slightly anterior to middle, more narrowed posteriad than anteriad, with anterior angles rounded, slightly protruded anteriad; median portion with or without moderately short and deep longitudinal impression in antero-median portion; postero-lateral portions distinctly impressed and reflexed. Punctation dense, coarse, and deep, sparser and finer along middle, sometimes with impunctate area in postero-median third.

Elytra slightly wider than long, slightly widened posteriad, with straight or rounded hind margins. Punctation similar to that of pronotum, sometimes sparser and somewhat finer in apical portion.

Metatarsus more than twice as long as metatibia.

Abdomen with two transverse, large tomentose spots in the middle of tergite IV, and slightly smaller spots in the middle of tergite V. Punctation distinct, dense, rather deep.

Male. Posterior margin of tergite VIII rounded. Posterior margin of sternite VIII sinuate. Aedeagus ( Fig. 6 ) with moderately wide median lobe, in apical portion gradually narrowed toward acute apex; parameres slightly extending beyond apex of median lobe, with three moderately short apical setae, medial margins without small setae; internal sac with long flagellum, slightly widened basally. Aedeagus laterally as in Fig. 7 .

Female. Posterior margins of tergite VIII and sternite VIII rounded.

Comparative notes. Regarding the wide median lobe of the aedeagus, G. spadiceus Shavrin , sp.n. is most similar to G. amplissimus , from which it differs by the narrower apical portion of the aedeagus, by the absence of additional setae on the medial edges of the narrower parameres, and by a slightly wider flagellum. The new species differs from all the species of the crassipalpis group by the reddish-brown elytra.

Etymology. The specific epithet is the Latin adjective spadiceus (reddish-brown). It refers to the coloration of the elytra.

Distribution. The species is at present known only from the type locality in the Manaslu range, Central Nepal ( Fig. 7 ).

Natural History. The specimens were collected at an elevation of 3900 m a.s.l. Detailed ecological data are unknown.

Figures

FIGURE 11. Distribution of Geodromicus crassipalpis (black circles) and G. spadiceus (black square) in the Himalayan Region.

FIGURES 1–3. Habitus of species of Geodromicus crassipalpis group: 1–G. crassipalpis (Uttarakhand), 2–G. amplissimus (holotype), 3–G. spadiceus (holotype). Scale bar: 1.0 mm.

FIGURES 4–10. Aedeagus of species of Geodromicus crassipalpis group (4–5: G. crassipalpis (Uttarakhand); 6–7: G. spadiceus (paratype); 8–9: G. amplissimus (paratype; Yunnan); 10: G. sp. (Gansu)): 4, 6, 8, 10–parameral view, 5, 7, 9–lateral view. Scale bar: 0.1 mm.