Cis incomptus Lawrence, 2019

Lawrence, John F., 2019, The Australian Ciidae (Coleoptera: Tenebrionoidea): supplement, Zootaxa 4555 (4), pp. 451-490 : 469-470

Cis incomptus Lawrence sp. nov.

( Figs 21–22 , 61, 79)

Diagnosis. This is a relatively small, setose species with dual and confused elytral punctation and no obvious frontoclypeal or pronotal modifications in the male. It is known from far northern QLD and the Torres Strait Islands, and may occur in New Guinea.

Description. With characters of the genus. Length = 1.00–1.20 (1.11 ± 0.06, n = 11) mm. Body moderately elongate and slightly convex. BL/EW 2.04–2.27 (2.14); GD/EW = 0.78–0.88 (0.82). Colour of head and pronotum reddish-brown to black, elytra yellowish-brown to black, undersides usually dark reddish-brown, legs yellowishbrown; antennal funicle yellow with club darker; surfaces shiny. Dorsal vestiture single, consisting of short, erect, yellow bristles, just visible under lower power. Frons and vertex flat in both sexes; without vertexal sex patch in male. Longest eye diameter 0.29 times as great as head width. Frontoclypeal ridge in both sexes complete at middle and slightly produced laterally. Antennomere length ratio: 5.25: 3.50: 3.00: 1.75: 1.00: 1.25: 1.00: 3.00: 3.75: 5.00; length/width ratios: 1.75, 1.56, 2.40, 1.17, 0,67, 0.71, 0.50 0.83, 0.94, 1.25. Apical maxillary palpomere 2.36 times as long as wide, widest at basal third, apex obliquely truncate. Labial palps separated by 0.17 times basal width of one; ratio of palpomere lengths 1.00: 2.67: 2.00, apical palpomere narrower than preapical one, which is inflated. Male gula 0.42 times as wide as head, densely filled with micropores, except near posterior edge, the pores extending anteriorly onto submentum as a moderately narrow, apically rounded patch; female gula 0.32 times as wide as head. Pronotum 0.79–0.91 (0.86) times as long as wide, widest at about middle in both sexes; anterior edge strongly rounded in both sexes; lateral margins very narrow, not visible for their entire lengths from above, with smooth edges; anterior angles not produced forward; posterior angles rounded; posterior edge with very narrow but complete marginal bead; disc evenly convex; punctation moderately fine but dense, punctures usually separated by less than a diameters; interspaces finely sculptured and somewhat shiny. Prosternum in front of coxae 1.14 times as long as mid length of procoxal cavity, slightly convex, prosternal process 0.71 times as wide as mid length of coxal cavity, truncate at apex; procoxal cavities very narrowly open; postcoxal process narrowly acute. Elytra 1.31–1.42 (1.36) times as long as wide and 1.58–1.85 (1.74) times as long as pronotum; punctation dual and confused, with megapunctures about the same size as pronotal punctures, the interspaces smooth and shiny. Mesocoxal cavities separated by about 0.14 times shortest diameter of a cavity. Metaventrite slightly convex; discrimen about 0.50 times length of ventrite; shortest distance between meso- and metacoxal cavities about 1.29 times length of abdominal ventrite 1. Outer apical angle of protibia in both sexes somewhat angulate. First abdominal ventrite 2.00 times as long as 2nd; sex patch in male about 0.43 times as long as ventrite 1, located slightly behind middle, circular, with distinct rim. Sides of sternite VIII straight and converging, its apex weakly emarginate, with more or less rounded angles. Pregenital ring relatively broad with acute apex. Tegmen (Fig. 61) 2.05 times as long as wide, widest at apical fifth, sides slightly converging to basal third, then gradually diverging to apical fifth where they converge to apical lobe, which is broad and somewhat truncate with a slight notch at middle; just before apical lobe on each side is a shelf bearing a short, rounded lobe and an articulated gonostylus-like process with a long seta at its apex; base subacute. Penis 1.10 times as long as tegmen and 4.0 times as long as wide, sides slightly converging to beyond middle, slightly diverging to apical fifth, then strongly diverging to acute apex; base broadly rounded. Ovipositor (Fig. 79) 1.30 times as long as wide, widest at base with sides subparallel but gradually converging at apical third; paraprocts as long as gonocoxites, longitudinally oriented; gonocoxites 0.60 times as long as their combined widths; proximal lobe slightly transverse, not subdivided; distal lobe about 2.5 times as long as proximal lobe gradually narrowed to just beyond middle, then abruptly narrowed forming a horizontal mesal shelf and a narrow, subcylindrical lateral lobe with subtruncate apex; a pair of ventrally projecting lobes just proximal to shelf; gonostylus 0.28 times as long as distal lobe and 3.5 times as long as wide.

Type specimens: Holotype, ♂, “Claudie R. nr. Iron Rg. Qld 19–25 July 1978 J. F. Lawrence / J. F. Lawrence Lot 78-97 Trametes scabrosa / ANIC Image” ( ANIC 25-014994 ).

Paratypes. QLD: Badu Island , Torres Strait, C. T. McNamara (4, SAM, ANIC) ; Claudie River , near Iron Range, 19–25.vii.1978, 78-97 JFL, Trametes scabrosa, J. F. Lawrence (5, ANIC) ; Claudie River , near Iron Range, 19–25.vii.1978, 78-74 JFL, Trichaptum sp., J. F. Lawrence (1, ANIC) ; West Claudie River , N. Iron Range (12.45S, 143.14E), 50m, 3.xii.1985, QM berlesate No. 690, stick brushing, rainforest, G. B. Monteith (1, QMB) .

Distribution. Known only from northern Cape York and Torres Strait Islands.

Biology. Feeding in basidiomes of Trametes scabrosa and Trichaptum sp. and found in berlesates from stick brushing.

Etymology. Derived from the Latin incomptus , meaning unadorned or simple and referring to the lack of obvious secondary sexual characters in this species.

Figures

FIGURES 16–30. habitus:, Cis depressus sp. nov.: 16, female, dorsal; 17, same, lateral; 18–20, Cis howensis sp. nov.: 18, male, dorsal (1.6 mm); 19, female, dorsal (1.6 mm); 20, same, lateral; 21–22, Cis incomptus sp. nov.: 21, male, dorsal (1.0 mm); 22, same, lateral; 23–25, Cis latemarginatus sp. nov.: 23, male, dorsal (1.1 mm); 24, same, lateral; 25, female, dorsal (1.1 mm); 26–28, Cis macilentus sp. nov.: 26, male, dorsal (1.3 mm); 27, female, dorsal (1.2 mm); 28, male, lateral; 29–30, Cis occidentalis sp. nov.: 29, male, dorsal (1.3 mm); 30, female, dorsal (1.3 mm).