SCANSORIOPTERYGIDAE

Pittman, Michael & Xu, Xing, 2020, Pennaraptoran Theropod Dinosaurs Past Progress And New Frontiers, Bulletin of the American Museum of Natural History 2020 (440), pp. 1-353 : 44-46

SCANSORIOPTERYGIDAE

Scansoriopterygids are a bizarre group of early-diverging Laurasian oviraptorosaurians or paravians, known only from the Middle and Late Jurassic Haifanggou Formation and Late Jurassic Tiaojishan Formation of north China so far (~168–155 Ma) (Czerkas and Yuan, 2002; Zhang et al., 2002; Zhang et al., 2008a; Turner et al., 2012; Brusatte et al., 2014; Xu et al., 2015 a; Wang et al. 2019 a; Pei et al., in press) (fig. 1; table 2). Known from five (or six: O’Connor and Sullivan, 2014) feathered Chinese specimens, only one definitive and possibly two somatically mature individuals exist. Two of these specimens (Yi qi and Ambopteryx longibrachium ) possess feathered, membranous wings ( Xu et al., 2015 a; Wang et al., 2019 a) and one possesses a pygostyle ( Wang et al., 2019 a). Epidendrosaurus and Epidexipteryx are two well-accepted genera, but Scansoriopteryx may be the same genus as Epidendrosaurus . The Early Cretaceous Zhongornis , originally described as a bird (Gao et al., 2008), may be a scansoriopterygid instead (O’Connor and Sullivan, 2014), but this has been contested (Rashid et al., 2018). The notion that scansoriopterygids are early-branching avialans (Xu et al., 2011a; Czerkas and Feduccia, 2014) has been replaced by anatomical evidence grouping some or all scansoriopterygids with oviraptorosaurians (Turner et al., 2012; Agnolín and Novas, 2013; Brusatte et al., 2014; Pei

TABLE 3

Dromaeosaurid fossil record

Geological Unit Country Period Age Age Reference Taxa Reference

Bayan Gobi Early

China Aptian-Albian Pittman et al., 2015 IVPP V22530 Pittman et al., 2015 Formation Cretaceous

Changyuraptor Xu et al., 1999 , 2000, 2003; Xu

Graciliraptor , and Wang, 2004a; Hwang et al., Jehol Group

He et al., 2004; Chang et Sinornithosaurus , 2002; Longrich and Currie, 2009; (Yixian Formation; Early Barremian-

China al., 2009, 2017; Pan et al., Tianyuraptor, Zheng et al., 2009 ; Gong et al., Jiufotang Cretaceous Aptian

2013 Zhenyuanlong , 2012; Han et al., 2014; Lü and Formation)

Zhongjianosaurus ; Brusatte, 2015; Xu and Qin, 2017;

Microraptor , Wulong Pei et al., 2014 ; Poust et al., 2020 Wulansuhai/ Bayan

Late Linheraptor , Velociraptor Godefroit et al., 2008 ; Xu et al., Mandahu China Campanian Godefroit et al., 2008

Cretaceous osmolskae 2010b, 2015a Formation

Late Campanian-

Qiupa Formation China Jiang et al., 2011 Luanchuanraptor Cretaceous Maastrichtian

Lü et al., 2007

Shuvalov, 2000; van

Bayan Shireh Late Cenomanian-

Mongolia Itterbeeck et al., 2005; Achillobator Perle et al., 1999 Formation Cretaceous Santonian

Kurumada et al., 2020

van Itterbeeck et al., Halszkaraptor, Osborn, 1924 ; Norell et al., Djadokhta Late

Mongolia Campanian 2005; Dingus et al., 2008; Mahakala , Tsaagan , 2006; Turner et al., 2007b, 2011; Formation Cretaceous

Hasegawa et al., 2009 Velociraptor mongoliensis Cau et al., 2017

Gradzinski and

Barun Goyot Late Campanian- Hulsanpes (possibly not Osmólska, 1982; Turner et al., Mongolia Jerzykiewicz, 1974a,

Formation Cretaceou Maastrichtian a dromaeosaurid) 2012; Cau and Madzia, 2018

1947b; Fanti et al., 2012

Jerzykiewicz and Russell,

Late

Nemegt Formation Mongolia Maastrichtian 1991; Shuvalov, 2000; van Adasaurus Barsbold, 1983

Cretaceous

Itterbeeck et al., 2005

Öösh Formation Mongolia Early Berriasian- Turner et al., 2007c Shanag Turner et al., 2007c

Cretaceous Barremian

South Early suspected

Jinju Formation Aptian Kim et al., 2018 Kim et al., 2018

Korea Cretaceous microraptorine tracks

Late Sues and Averianov, Kurzanov, 1976; Sues and Bissekty Formation Uzbekistan Turonian Itemirus

Cretaceous 2014, 2015 Averianov, 2014

3 continued 3 continued et al., in press) or as early-branching paravians (Turner et al., 2012; Godefroit et al., 2013a, 2013b; Xu et al., 2015 a; Wang et al., 2019 a).