Thyestetha carbonaria Faust

Eberle, Jonas, Tänzler, Rene & Riedel, Alexander, 2012, Revision and phylogenetic analysis of the Papuan weevil genus Thyestetha Pascoe (Coleoptera, Curculionidae, Cryptorhynchinae), Zootaxa 3355 (1), pp. 1-28 : 5-6

Thyestetha carbonaria Faust

( Figs. 1 , 10 , 26, 37–38, 43, 66, 71)

Thyestetha carbonaria Faust 1898: 194 f. (type locality: Fergusson Isl.).

Diagnosis. Pronotum densely punctate with small punctures, black. Elytra dark ferruginous, almost black, with distinct small punctures, interstices 2– 4 X diameter of punctures ( Figs. 1 , 10 ). Rostral canal with scattered white scales. Dorsal squamose line of metafemur moderately dense. Aedeagus ( Figs. 37–38 ) with apex strongly curved ventrad, medially slightly extended ( Fig. 43 ).

Redescription. Lectotype, male. Length. Pronotum + elytra: 4.6 mm. Color. Antenna, rostrum and tarsi ferruginous; pronotum black; head, elytra, pleural sclerites, legs and ventral surface dark ferruginous, almost black. Habitus subovate, in dorsal view almost without constriction between pronotum and elytra ( Fig. 1 ); profile dorsally with shallow constriction between pronotum and elytron ( Fig. 10 ).

Rostrum 5.6 X as long as wide at base; base coarsely punctate, base dorsally with scattered scales. Head. Vertex coarsely punctate, few white scales near eye.

Prothorax 1.2 X wider than long; disc uniformly densely punctate with small punctures, basally slightly sparser, punctures slightly smaller near pronotal base (in front of basal row of coarse punctures), interstices smooth, polished; sides densely punctate with larger punctures; postocular lobe foveate-rugose; marginal carina externally along edge with few whitish scales; rostral canal with scattered long setae and between procoxae with whitish scales.

Elytra as long as wide; sides rounded; apex slightly extended, jointly rounded; punctures of striae 1–9 distinctly impressed from base to apex; interstices between punctures 2– 4 X wider than diameter of punctures, punctures closer to each other apically, interstices between punctures of stria 9 ca. 0.5 X diameter of puncture. Metanepisternum 2.8 X wider than high.

Thoracic venter. Meso- and metaventrite laterally with sparse whitish scales; metaventrite submedially behind mesocoxa with pair of moderate protrusions. Pro- and mesocoxa densely punctate, squamose; metacoxa with scales near femoral insertion; trochanter with sparse, coarse punctures containing small scales. Distance between mesocoxae subequal to width of mesocoxa.

Legs. Femora moderately punctate, scales laterally slender, dorsally broader; metafemur with denser scaling, dorsal squamose line moderately dense; anteroventral ridges weakly developed; with posteroventral ridge in apical third abruptly ending and markedly protruding proximally (Fig. 26).

Abdomen. Ventrite 1 and 2 coarsely punctate, laterally squamose, anterior margin of ventrite 2 smooth; remaining ventrites subglabrous.

Terminalia. Aedeagus ( Figs. 37, 38, 43 ). Apex rounded, strongly curved ventrad, sides weakly sinuate; lateral orificial sclerites medium-sized, inner portion ca. 2 X longer than outer portion; anterior median orificial sclerite simple between 2nd and 3rd fold (as in Fig. 36 ). Dorsal contour of transfer apparatus sharply incised at middle, ventral contour angularly projecting; sclerites in basal half, dorsoventrally flattened, moderately distant, in apical half contiguous. Sclerotized part of ventral membrane extending proximad 0.82 X length of aedeagal body from insertion of apodemes.

Female paralectotype. Same as lectotype except: Rostrum 5.60 X longer than wide. Prothorax 1.26 X longer than wide. Elytra 1.04 X longer than wide; extreme apex slightly more slender. Metafemur without concavity basal to stridulatory patch, posteroventral ridge not protruding.

Intraspecific variation. Size: Length 4.5–4.6 mm (n=2, x = 4.6 mm) in males. White squamose line on dorsal surface of metafemora more or less distinct.

Material examined (2 males, 1 female). Male, lectotype by present designation: PAPUA NEW GUINEA, Milne Bay Prov., Fergusson Island ; labels ( Fig. 66 ): 1) [golden rectangle] 2) [rectangular, white] Fergusson I., ix. x. xi. xii. 9 (A. S. Meek) [printed, with male gender symbol hand-written on the left] 3) [rectangular, white] carbonaria Faust [hand-written] 4) [rectangular, blue] Coll. J. Faust, Ankauf 1900 [printed] 5) [rectangular, red] Type [printed] 6) [rectangular, white] Staatliches Museum für Tierkunde Dresden [printed] 7) [rectangular, red] LECTOTYPE, Thyestetha carbonaria Faust [printed] condition: onychium of right protarsus missing; right mesotarsus broken, glued on card; specimen originally point-mounted, remounted on card; genitalia in glycerol in microtube ( SMTD) . Paralectotype: 1 female: PAPUA NEW GUINEA, Milne Bay Prov., Fergusson Island; labels: 1) [golden rectangle] 2) [rectangular, white] Fergusson I., ix. x. xi. xii. 95 (A. S. Meek) [printed] 3) [rectangular, white] carbonaria Faust [hand-written] 4) [rectangular, blue] Coll. J. Faust, Ankauf 1900 [printed] 5) [rectangular, red] Type [printed] 6) [rectangular, white] Staatliches Museum für Tierkunde Dresden [printed] 7) [rectangular, red] PARALECTOTYPE, Thyestetha carbonaria Faust, 1898 , [printed] in perfect condition, glued on original card ( SMTD).

Other specimen: Male: PAPUA NEW GUINEA, Milne Bay Province, Fergusson Island ( SMTD) .

Distribution ( Fig. 71 ). Milne Bay Prov. (Fergusson Isl.).

Notes. For this study, three syntypes of T. carbonaria from Milne Bay and Fergusson Island were available. One of them belongs to a different species described below as T. oblita sp. n., Therefore, a lectotype is designated to ensure stability of nomenclature. FAUST (1898) did not specify the exact number of specimens examined. One or more paralectotypes from the Trobriand Islands, we were unable to locate.

Figures

FIGURE 71. Distribution of Thyestetha carbonaria, T. discolor sp. n., T. emarginata sp. n., T. nitida, and T. oblita sp. n..

FIGURES 1 – 9. Habitus of Thyestetha spp., dorsal aspect: (1) T. carbonaria, lectotype; (2) T. discolor sp. n., holotype; (3) T. emarginata sp. n., holotype; (4) T. glabra sp. n., holotype; (5) T. lubrica sp. n., holotype; (6) T. nitida; (7) T. oblita sp. n., holotype; (8) T. puncticollis sp. n., holotype; (9) T. splendens sp. n., holotype.

FIGURES 10 – 18. Habitus of Thyestetha spp., lateral aspect: (10) T. carbonaria, lectotype; (11) T. discolor sp. n., holotype; (12) T. emarginata sp. n., holotype; (13) T. glabra sp. n., holotype; (14) T. lubrica sp. n., holotype; (15) T. nitida, lectotype; (16) T. oblita sp. n., holotype; (17) T. puncticollis sp. n., holotype; (18) T. splendens sp. n., holotype.

FIGURES 37 – 54. Aedeagus of Thyestetha spp. in dorsal and lateral aspect: (37, 38) T. carbonaria, lectotype; (39, 40) T. discolor sp. n., holotype; (41, 42) T. glabra sp. n., holotype; (43) T. carbonaria, apex of aedeagus, ventral aspect; (44) T. oblita sp. n., holotype, apex of aedeagus, ventral aspect; (45, 46) T. lubrica sp. n., holotype; (47, 48) T. nitida, lectotype; (49, 50) T. oblita sp. n., holotype; (51, 52) T. puncticollis sp. n., holotype and (53, 54) T. splendens sp. n., holotype.

FIGURES 29 – 36. Male terminalia of Thyestetha spp.: (29 – 31) T. glabra sp. n., holotype: (29) tegmen, (30) tergite VII and (31) segments VIII – IX. (32 – 34) T. nitida: (32) tegmen, (33) tergite VII and (34) tergite VIII with sternite VIII and spiculum gastrale. Ostium, lateral aspect: (35) T. discolor sp. n., arrow and dashed line marking bulge between 2 nd and 3 rd fold (36) T. splendens sp. n..

FIGURES 66 – 67. Collection labels of Thyestetha spp.: (66) T. carbonaria, lectotype; (67) T. nitida, lectotype. FIGURES 68 – 70. Pantoxistus rubripennis, lectotype: (68) collection labels; (69) dorsal aspect, (70) ventral aspect.