Neobarombiella flavilabris ( Weise, 1903 ) Bolz & Wagner, 2012

Bolz, Helmut & Wagner, Thomas, 2012, 3463, Zootaxa 3463, pp. 1-112 : 14-20

Neobarombiella flavilabris ( Weise, 1903) comb. n.

( Figs 2–23)

Monolepta flavilabris Weise, 1903: 213 .

Candezea acutangula Weise, 1903: 215 syn. n.

Candezea vivida Weise, 1909: 211 syn. n.

Barombia benguelensis Laboissière, 1921: 287 syn. n.

Barombia gregaria Laboissière, 1931: 51 syn. n.

Barombiella bifasciata Laboissière, 1939: 159 syn. n.

Barombiella clypeata Laboissière, 1940: 82 syn. n.

Barombiella ghesquierei Laboissière, 1940: 82 syn. n.

Barombiella trifasciata Laboissière, 1940: 84 syn. n.

Barombiella vicina Laboissière, 1940: 83 syn. n.

Barombiella wittei Laboissière, 1940: 85 syn. n.

Barombiella alluaudi Laboissière , in litteris

Total length. 3.45–5.50 mm (mean: 4.41 mm) (n=36).

Head. Frons and vertex either entirely yellow (about 10% of material examined), reddish-yellow (40%), or black (50%; Figs 7 a–f), head often darker than pronotum. Labial and maxillary palpi yellow to brownish-black. Antenna pale yellow to yellow, often becoming more brownish towards apex, last antennomere usually dark brown to black ( Fig. 7 a–f). Length of second to third antennomere 0.67–0.78 (mean: 0.72), length of third to fourth antennomere 0.62–0.73 (mean: 0.68). Eyes large and with short interocular distance ( Figs 7 a–f), width of eye to interocular distance 0.60–0.69 (mean: 0.64).

Thorax. Pronotum yellow in 20% of material examined ( Fig. 7a), all other specimens yellowish-red to red ( Figs 7 b–f), about half of specimens with brownish to black spots in outer posterior corners ( Figs 7d, f); broad, strongly trapezoidal, distinctly and deeply punctuated and with distinct micro-sculpture. Pronotal width 1.30–2.06 mm (mean: 1.71 mm), pronotal length 0.62–0.96 mm (mean: 0.78 mm), pronotal length to width 0.43–0.49 (mean: 0.46). Elytron rarely entirely yellow (10%), if so, whole specimen usually yellow or even pale yellow, as in lectotype of Candezea acutangula ( Fig. 13b); other 90% of specimens with black humeral spots and in nearly all these specimens, with transverse brown or black band in apical third ( Figs 7 b–f, 12b, 14b–15b, 17b – 23b) and very rarely with predominantly black elytron and yellow spots; deeply and densely punctuated with indistinct micro-sculpture; elytral length 2.70–4.25 mm (mean: 3.50 mm), elytral width 1.90–2.80 mm (mean: 2.44 mm), maximal width of both elytra to length of elytron 0.66–0.75 (mean: 0.71). Meso- and metathorax yellow, rarely brownish. Legs entirely yellow, length of basi-metatarsi to metatibiae 0.44–0.52 (mean: 0.48).

Abdomen. Yellow.

Male genitalia. Median lobe slender, slightly conical from middle towards apex which is distinctly downcurved ( Figs 8 a–c); apical section lacking ridges or spikes, the apex rounded in lateral view and poorly sclerotized; endophallic brush with two delicate, slender spiculae covered by median lobe and tectum.

Diagnosis. Most similar to Neobarombiella senegalensis . Both species can be distinguished by the shape of their antennomeres, which are slender. They are broader and shorter in N. senegalensis . The third antennomere in N. flavilabris is shorter than the fourth (length of third to fourth antennomere: 0.62–0.73; N. senegalensis 0.47–0.54; Figs 9, 93). The elytron of N. flavilabris are more densely and coarsely punctuated than the usually finely punctuated dorsum in N. senegalensis . However, since these external characters might be atypical for some specimens, only the male genitalia allow distinct differentiation of both species ( Figs 8, 93). The median lobe of N. flavilabris is very short and down-curved apically, with a poorly sclerotized apex and no ridges, while N. senegalensis has a median lobe which is more straight, with a beak-like apical section and two strongly sclerotized ventral ridges.

Other similar species are N. nigrosuturalis sp. n. and N. frontalis sp. n. which can be distinguished both by their black sutural margins ( Figs 7, 153, 173), N. cornuta sp. n. and N. medvedevi sp. n. by their short second antennomere (length of second to third antennomere: 0.67–0.78; 0.55–0.60 in N. cornuta sp. n.; 0.58–0.64 in N. medvedevi sp. n.; Figs 139, 167). Neobarombiella naumanni sp. n. has smaller eyes and a larger interocular distance, width of eye to interocular distance 0.50–0.57 in N. naumanni sp. n. and 0.60–0.69 in N. flavilabris ( Figs 7, 169), and N. fassbenderi sp. n. is much smaller having a total length of 2.65–3.40 mm compared to 3.45–5.50 mm in N. flavilabris ( Figs 7, 145)

Remarks. A wide distribution, obvious high abundance in tropical Africa, and particularly high degree of variability in dorsal colouring has led to no less than ten synonymies in this species. Laboissière, especially in his last publications in 1939 and 1940, described it several times based on distinct colour patterns. Based on the hundreds of specimens revised in this study, it is clear that there are no distinct types of colouring, but rather a continuous polychromatism from entirely pale yellow, over many specimens with more or less broad transverse elytral bands to predominantly black elytron.

Distribution. Nearly pan-Afrotropical, from Sierra Leone to Ethiopia, southwards through the Congo Basin towards Angola and northern and eastern South Africa ( Fig. 11).

Type material examined.

Monolepta flavilabris . Holotype, male: “Type / Pangani Steppe / Pangani Steppe / Monolepta , flavilabris, m. / ex coll., J. Weise / Holotypus, Monolepta , flavilabris Wse, 1903 / Neobarombiella flavilabris, ( Weise, 1903) , Bolz & Wagner det. 04 / AfriGa, specimen ID, 1031, specimen data, documented, 20.VII.2005 “ ( MNHU; Fig. 12). There is only one specimen available from the type locality, which can be treated as holotype. Colouring type Fig. 7c. Type locality: Tanzania, Pangani, 5°26'S / 38°58'E .

Candezea acutangula . Lectotype, female: “Type / Nguelo / acutangula, m. / ex. coll., J. Weise / Type / Lectotypus, Th. Wagner desig. 2005, Candezea acutangula Wse. 1903 / Neobarombiella flavilabris, ( Weise, 1903) , Bolz & Wagner det. 04 / AfriGa, specimen ID, 952 specimen data, documented, 5.VII.2005 ” ( MNHU; Fig. 13). Weise listed three localities in Tanzania “Nguelo, Mombo, and Kwai”, and gave no details on specimen numbers. A lectotype is here designated to fix the species name on a single specimen; same for all other lectotype designations in this paper. Colouration type Fig. 7a. Type locality: Tanzania, Nguela, 4°45'S / 38°30'E .— Paralectotypes. 3 ex., Usambara , Nguela, 4°45'S / 38°30'E ( MNHU) ; 1 ex. Mombo , 4°53’S / 38°17’E, VII .1899, ex coll. J. Weise (MNHU); 1 ex., Kwai , 4°44’S / 38°21’E, Paul, ex coll. J. Weise ( MNHU) .

Candezea vivida . Holotype, female: “Kilimand., Sjöstedt / Kilimandjaro, Sjöstedt 1905-6 / Kibonoto, 1300–1900 m. / Type / Candezea , vivida m / ex coll., J. Weise / Holotypus, Candezea , vivida Weise, 1909 / Neobarombiella flavilabris, ( Weise, 1903) , Bolz & Wagner det. 04 / AfriGa, specimen ID, 1153, specimen data, documented, 5.VII.2005 ” (MNHU; Fig. 14). Weise listed only one specimen “ 1 Exemplar ... Kulturzone des Kilimandjaro bei Kibonoto am 27. April” which can be treated as holotype by original designation. Colouring type like Fig. 7c, but with yellow head and pronotum. Type locality: Tanzania, Kilimandjaro, Kiboto, 3°09’S / 36°51’E.

Barombia benguelensis . Holotype, female: “Muséum Paris, Angola, Capelongo près Lucequé, (Benguela) Haut Cunene, Mission Rohan-Chabot, 1914 / Décembre / Barombia , benguelensis, m., V. Laboissière – Dét. / Type / Neobarombiella flavilabris ( Weise, 1903) Bolz & Wagner det. 04 / AfriGa, specimen ID, 1205, specimen data, documented, 20.IX.2004 ” (last cited label not included with figure; MNHN; Fig. 15). There is no specific information on type specimens in the original description. Since there is only one locality and date given, this might be the only one. The specimen also bears a separate “Type” label in Laboissière’s hand-writing and is here treated as the holotype. Colouration type similar to Fig. 7c but with accentuated black elytral colouration. Type locality: Angola, Capelongo near Lucequé , 14°39'S / 15°4'E.

Barombia gregaria . Lectotype, female: “Muséum Paris, Mozambique, Haut Nhamapaza, Forèt, S. de Maringoué, P. Lesne 1929 / Juin / Type / Barombia , gregaria, m., V. Laboissière – Dét. / AfriGa, specimen ID, 1217, specimen data, documented, 20.IX.2004 ” ( MNHN; Fig. 16). Laboissière listed “un seul individu” from Nova Choupanga près Chemba, and “en grand nombre” from Foret au sud de Maringoué dans le Haut Nhampaza. There are 31 type specimens available and the lectotype is here designated. Colouring type Fig. 7a. Type locality: Mozambique, Haut Nhamapaza, 18°02’S / 35°36’E .— Paralectotypes: the rest of the specimens in the series are paralectotypes.

Barombiella bifasciata . Lectotype, female: “Cotype / Paratype, bifasciata / Musée du Congo, Elisabethville, X-1911, Miss. Agric. / Barombiella , bifasciata m., V. Laboissière – Dét. / R. Dét., F, 3640 / Lectotypus, Th. Wagner desig. 2005, Barombiella , bifasciata Lab. 1939 / Neobarombiella flavilabris, ( Weise, 1903) , Bolz & Wagner det. 04 / AfriGa, specimen ID, 1207, specimen data, documented, 5.VII.2005 ” ( MRAC; Fig. 17). Laboissière listed four specimens from three localities without designation of a holotype. A lectotype is here designated. Colouring type Fig. 7e. Type locality: Democratic Republic of the Congo, Elisabethville, 11°40'S / 27°28'E .— Paralectotypes: the three remaining specimens are paralectotypes.

Barombiella clypeata . Holotype, female: “ Type / Musée du Congo, Uelé: Monga, 18–iv–8–v–1935, G. F. de Witte, Parc Nat. Albert / Barombiella , clypeata, m., V. Laboissière – Dét. / B. Type, clypeata / R. Dét., V, 4572 / Holotypus, Barombiella , clypeata Lab., 1940 / Neobarombiella flavilabris, ( Weise, 1903) , Bolz & Wagner det. 04 / AfriGa, specimen ID, 1209, specimen data, documented, 5.VII.2005 “ ( MRAC; Fig. 18) . Holotype by monotypy as indicated in the original publication. Colouring type similar Fig. 7c but with enhanced black elytral colouring. Type locality: Democratic Republic of the Congo, Uelé, Monga, 4°12’N / 22°49’E .

Barombiella ghesquieri . Lectotype, male: “Type / Musee du Congo, Rutshuru, – V–1937, J. Ghesquière, 4503 / Barombiella , ghesquierei, m, V. Laboissière – Dét. / TYPE, ghesquierei / R., Dét., S, 3921 / Lectotypus, Th. Wagner desig. 2005, Barombiella , ghesquieri Lab. / Neobarombiella flavilabris, ( Weise, 1903) , Bolz & Wagner det. 04 / AfriGa, specimen ID, 1214, specimen data, documented, 5.VII.2005 ” ( MRAC; Fig. 19). Laboissière indicated at least four type specimens from Rutshuru “ 4 spécimens, 2 typiques et 1 dans chaque variété”, a lectotype is here designated. Colouring type similar to Fig. 7c but posterior transverse elytral band reduced to large black subcircular spots, each one on each elytron. Type locality: Democratic Republic of the Congo, Rutshuru, 1°11’S / 29°27’E .— Paralectotypes: four specimens from the same location as the lectotype ( MRAC) .

Barombiella trifasciata . Holotype, male: “ Type / Musée du Congo, Camp Ruindi (1000), 20/ 28–XI–1934, G.F. de Witte, Parc Nat. Albert, 828 / Barombiella , trifasciata m., V. Laboissère – Dét. / B. Type, trifasciata / R. Det., T, 4572 / Holotypus, Barombiella , trifasciata Lab., 1940 / Neobarombiella flavilabris, ( Weise, 1903) , Bolz & Wagner det. 04 / AfriGa, specimen ID, 1248, specimen data, documented, 5.VII.2005 ” ( MRAC; Fig. 20) . Holotype by monotypy. Colouring type similar to Fig. 7c. Type locality: Democratic Republic of the Congo, Rwindi, 0°48’S / 29°18’E .

Barombiella vicina . Holotype, female: “ Holotypus / Type / Musée du Congo, Ndeko près Rwindi, (1083) 27- XI-1934, G.F. de Witte, Parc Nat. Albert, 820. / Barombiella , vicina m., V. Laboissière – Dét. / R. Det., X, 4572 / Type, B. vicina / Holotypus (covered on Fig. 21a: Barombiella vicina Lab. 1940 ) / Neobarombiella flavilabris, ( Weise, 1903) , Bolz & Wagner det. 04 / AfriGa, specimen ID, 1258, specimen data, documented, 5.VII.2005 ” ( MRAC; Fig. 21) . Holotype by original designation. Colouring type similar Fig. 7c but with yellow pronotum. Type locality: Democratic Republic of the Congo, Ndeko near Rwindi, 0°45’S / 29°20’E.— one specimen, Lemba, 1.–15.IV.1934, R. Mayné, is a paratype ( IRSN) .

Barombiella wittei . Holotype, female: “ Type / Congo belge: P.N.A., Ndeko (près Rwindi), 1082m. 27.xi.1934, G. F. de Witte: 835 / Barombiella , wittei m., V. Laboissière—Dét. / Coll. Mus. Congo / Type, B. wittei / R. Dét., W, 4572 / Holotypus / Holotypus, Barombiella , wittei Lab. 1940 / Neobarombiella flavilabris, ( Weise, 1903) , Bolz & Wagner det. 04 / AfriGa, specimen ID, 1260, specimen data, documented, 5.VII.2005 ” ( MRAC; Fig. 22) . Holotype by monotypy. Colouring similar to Fig. 7c but with yellow pronotum and the elytral suture broadly reddish in colour. Type locality: Democratic Republic of the Congo, Ndeko near Rwindi, 0°45’S / 29°20’E .

Further material with type labels, but unpublished

Barombiella alluaudi . female: “ Museum Paris / Afrique Orient. All., Kilimandjaro, Kiboscho, Ch. Alluaud, 1904 / Mars / Type / Barombia , alluaudi Labois, V. Laboissière – Dét. / Neobarombiella flavilabris ( Weise, 1903) , Bolz & Wagner det. 04” ( MNHN; Fig. 23). To our knowledge, a description of this species name has not been published, and it is hence unavailable within the meaning of the Code .

Other material examined. Cameroon. 1 ex., Muanamala, Dr. Chr. Schröder ( MNHU); 1 ex., Joko , 5°29'N / 12°19'E ( MNHU); 1 ex., Oyak II, 3°28'N / 11°30'E, IX.1970, B. de Miré ( MNHN); 1 ex., Pitoa, 9°23'N / 13°32'E, IX.1974, B. de Miré ( MNHN); 1 ex., Tibati, 6°28'N / 12°37'E, L. Colin ( MNHU) . – Democratic Republic of the Congo. 2 ex., Bas-Congo, Mayidi, 5°11'S / 15°09'E, 1952, Rév. P. van Eyen ( MRAC); 2 ex., Haut-Congo , 1897, Dr. Védy ( MRAC); 1 ex., Haut-Uele , Abimba , 4°20'S / 27°10'E, VII .1925, L. Burgeon ( MRAC); 1 ex., Ituri, Aru, 2°52'N / 30°51'E, 1938, Bastiaens ( MRAC); 1 ex., Katanga , Kolwezi , 10°50'S / 25°44'E, 1962, Dr. V . Allard ( MRAC); 1 ex., M'Paka, Libenge , 3°39'N / 18°38'E, VII .–VIII .1959, M. Pecheur ( MRAC); 5 ex., P. N. Garamba, 3°40'N / 29°00'E, VI .1950– III .1952 , G. Demoulin, H. de Saeger (IRSN); 2 ex., P. N. Upemba, 9°20'S / 27°12'E, XI.1947 – III .1949 , G. F. de Witte (IRSN); 40 ex., Elizabethville , 11°40'S / 27°28'E, VI.1949 – V .1959 , Ch. Seydel (MRAC); 1 ex., Kapiri, 9°41'S / 27°13'E, X.1912, Miss. Agric. (MRAC); 4 ex., Mayidi, 5°11'S / 15°09'E, 1942, Rév. P. van Eyen (MRAC); 43 ex., Musosa, 8°22'S / 29°37'E, X.1939, H. J. Brédo (MRAC); 1 ex., Ngowa , 5°42'S / 16°35'E, IV.1939, R. P. J. Mertens ( MRAC); 1 ex., Rabai , 3°55'S / 39°34'E, VIII .1937, van Someren ( NMK); 16 ex., Rutshuru , 1°11'S / 29°27'E, II.– V .1937, J. Ghesquiere ( MRAC); 2 ex., Sambasi, 4°10'S / 17°19'E, X.1906, F. Seiner ( MNHU); 1 ex., Stanleyville , 0°33'N / 25°14'E, IX.1928, A. Collart ( IRSN); 2 ex., XII .1912 ( IRSN); 1 ex., Brazzaville, 4°15'S / 15°17'E, 1904, Dr. J. Decorse ( MNHN); 5 ex., Congo da Lemba , 5°42'S / 13°42'E, V .1911– III.1913, R. Mayné ( MRAC); 1 ex., Costermansville, VII .1937, H. J. Brédo ( MRAC); 1 ex., Ubangi, Gemena, 3°15'N / 19°46'E, 1937, C. Léontovitch ( MRAC); 1 ex., Lulua , Kabomba , 4°31'S / 28°8'E, XI.1937, Vanderstichele ( MRAC); 1 ex., Lulua , Kapanga , 6°48'S / 22°32'E, XII .1932, G. F. Overlaet ( MRAC); 1 ex., Kisantu , 5°7'S / 15°5'E, XII .1920, Dr. H. Schouteden ( MRAC); 1 ex., Niembe, Lembe , 3°46'S / 13°19'E, VII .1918, R. Mayné ( MRAC); 1 ex., Lubumbashi , 11°40'S / 27°28'E, III .1975, W. Beun ( MRAC); 1 ex., Kivu, Lulenga, 0°37'S / 18°23'E, IX.1932, L. Burgeon ( MRAC); 1 ex., Kivu , Luvungi , 2°52'S / 29°2'E, XII .1932, L. Burgeon ( MRAC); 1 ex., Mpese 5°14'S / 15°33'E, VI .1937, R. P. J. Cooreman ( IRSN); 1 ex., P. N. Albert, Mutsora, 0°19'S / 29°45'E, 1939, Hackars ( MRAC); 1 ex., Nyangwe , 4°13'S / 26°11'E, III .1918, R. Mayné ( MRAC); 1 ex., Lac Kivu , Rwankwi, V .1950, J. V. Leroy ( MRAC); 1 ex., Wombali, 3°16'S / 17°20'E, IX.1913, P. Vanderijst ( MRAC); 1 ex., Haut-Congo, Yebo , 3°7'N / 24°39'E, XI.1925, L. Burgeon ( MRAC); 2 ex., P. N. Albert, 0°45'S / 29°20'E, XII .1956– I.1958, P. Vanshuytbroeck ( IRSN); 1 ex., Riv. Kabarazo, V .1936, L. Lippens ( MRAC) .— Equatorial Guinea. 48 ex., 2°00’N / 10°00’E, Dr. L. Báguena; 1 ex., Nniefang, 01°50’N / 10°08’E, Dr. L. Báguena ( MNCN); 1 ex., Valle del Mvulu, Nniefang, 01°51’N / 09°45’E, Dr. L. Báguena ( MNCN); 1 ex., Fernando Póo, Santa Isabel , 03°21’N / 08°40’E, VII .1919, Escalera ( MNCN); 8 ex., Fernando Póo, 03°35’N / 08°45’E, Dr. L. Báguena ( MNCN); 21 ex., Fernando Póo, Santa Isabel , 03°21’N / 08°40’E, Dr. L. Báguena ( MNCN); 1 ex., Fernando Póo , Basupú , 03°44’N / 08°49’E, V .1919, Escalera ( MNCN); 1 ex., Fernando Póo, Rebola, 03°43’N / 08°50’E, Dr. L. Báguena ( MNCN); 1 ex., Evinayong, 01°06’N / 10°17’E, Dr. L. Báguena ( MNCN); 1 ex., Alto Benito , Evinayong , 01°06’N / 10°17’E, Dr. L. Báguena ( MNCN) .— Eritrea. 1 ex., Decamere, V .1963, Linnavuori ( MZHF); 1 ex., Ghinda , 15°26'N / 39°5'E ( NHRS) .— Ethiopia. 3 ex., Lac Margarita, 1934–1935, R . de Meulenaere ( IRSN) .— Ghana. 1 ex., Abétifi, 6°40'N / 0°45'W ( MNHN); 1 ex., Aburi , 5°02'N / 1°46'W, 1912–1913, W. H. Patterson ( NMK) .— Ivory Coast. 2 ex., Bingerville, VIII .1962– II.1964, J. Decelle ( MRAC); 1 ex., Daloa , 6°52'N / 6°27'W, XII .1962, J. Decelle ( MRAC); 1 ex., Haut Sassandra, 4°58'N / 6°08'W, 1910, A. Chevalier ( MNHN); 1 ex., Mouyassue , 8°55'N / 7°25'W, XI.1961, J. Decelle ( MRAC); 1 ex., 1897, Delafosse ( MNHN) .— Kenya. 3 ex., Mt. Elgon, 1°7'N / 34°35'E, I.1992, O. Merkl & G. Várkonyi ( HNHM); 2 ex., Tenvo Nat. Park, Galana River , IV.1969, M. P. Clifton ( NMK); 2 ex., Chyulu Hills, 2°35'S / 37°50'E, IV.1938 ( NMK); 1 ex., Res. Mchelelo, 1°52'S / 40°8'E, XI.2001, J. Mugambi / W. Odero ( NMK); 2 ex., Tsavo, 2°59'S / 38°28'E, III .2000, Mauser ( NMK); 2 ex., Voi ( Tsavo ), 3°23'S / 38°34'E, XI.1996, M. Snížek ( MRSN); 1 ex. , Kakamega Forrest , 0°15'N / 34°52'E, I.1972, C. F. Huggins ( BMNH); 8 ex. , Kitui, 1°22'S / 38°01'E, X.1937, R . Toker ( NMK); 1 ex. , Machakos, 1°31'S / 37°16E, XI.1923, Dr. van Someren ( MNHU); 1 ex. , Meru, 0°03'N / 37°38'E, VII .1943, van Someren ( NMK); 6 ex., Nairobi , 1°17'S / 36°50'E, VIII .–X.1969, M.P. Clifton ( NMK); 1 ex., Rabai, 3°55'S / 39°34'E, I.1928, A. F. J. Gedye ( BMNH); 4 ex., Thika , 1°03'S / 37°05'E, X.1937, S. Blom-Bjorner ( BMNH) .— Liberia. 1 ex., Suring , I.1891, A. F. Demery ( RMNH) .— Malawi. 1 ex., Livingstonia , 10°36'S / 34°7'E, VII .2000, L. Schmidt ( MNHU) .— Mali. 1 ex., Sofara , 14°1'N / 4°14'W, 1909, R . Chudeau ( MNHN) .— Mozambique. 1 ex., Genge , 18°07'S / 35°52'E, III .1929, A. Collart ( ZMUH) .— Nigeria. 1 ex., Azare, 10°26'N / 12°07'E, 1928–1929, Dr. Lloyd ( BMNH); 1 ex., Ibadan, 7°23'N / 3°56'E, 1932, Clifford King ( MNHN); 1 ex., Mokwa , 9°16'N / 5°02'E, VIII .1970, P. H. Ward ( BMNH); 4 ex., S. Apeji , 6°58'N / 4°07'E, III .1956 ( BMNH); 2 ex., S. Apeji, 6°58'N / 4°07'E, 1965 ( BMNH); 1 ex., Yankari , 9°01'N / 12°19'E, XI.1984, V . Assing (CBe) .— Rwanda. 1 ex., Kinazi, I.1953, P. Basilewsky ( MRAC); 4 ex., Mahembe , 1°53'S / 29°31'E, I.1953, P. Basilewsky ( MRAC) .— Sierra Leone. 1 ex., Port Lokko , 8°46'N / 12°46'W, VIII .1925, E. Hargreaves ( BMNH) .— Somalia. 1 ex., Gesira, VI .1984, S. B. S. (Florenz); 1 ex., V . Duca Abruzzi ( MCSG) .— South Africa, ECape: 1 ex., Lusikisiki , 31°21'S / 29°35'E, II.1991, Richter ( MRSN) ; South Africa, KwaZulu-Natal. 2 ex., Epangeni , 28°44'S / 31°54'E, III .–X.1977, P. E. Reavell (1 ex. SANC, 1 ex. TMSA); 1 ex., Zululand, Eshowe , 28°53'S / 31°28'E, V .1926, R. E. Turner ( BMNH); 1 ex., Estcourt, XI.1892, G. A. K. Marshall ( BMNH); 1 ex., Giants Castle Res., 29°08'S / 25°27'E, I.1982, P. Reavell ( SANC); 1 ex., Gingindhlovu , 29°01'S / 31°34'E, V .1926, R. E. Turner ( BMNH); 1 ex., Injasuti, 29°8'S / 25°27'E, I.1982, P. Reavell ( SANC); 1 ex., Malvern, 29°52'S / 33°55'E, IX.1897, G. A. K. Marshall ( BMNH); 1 ex., Oribi Gorge Nat. Res. , 30°43'S / 30°16'E, I.2006, C. Chaboo & E. Grobbelaar ( SANC) ; South Africa, Limpopo. 1 ex., Letaba, 23°54’S / 30°04’E, XI.1981, Klapperich ( MRSN); 2 ex., Levubu Distr. , 23°04'S / 30°13'E, IV.1986, E. v. d. Linde ( SANC); 1 ex., Makuvha, 22°52'S / 30°29'E, II.1994, E. Grobbelaar ( SANC); 1 ex., New Agatha State, 23°57'S / 30°7'E, I.1991, B. Grobbelaar ( SANC); 1 ex., Nylstroom, 24°45'S / 28°15'E, IV.1919, Ludwig Pretorius ( SANC); 1 ex., Zoutpansberg, XII .1901, Magn. Knothe S. ( MNHU) .— Sudan. 1 ex., Equatoria, III .1963, Linnavuori ( MZHF); 1 ex., Malakal , 9°32'N / 31°39'E, L. Burgeon ( MRAC) .— Tanzania. 1 ex. , Arusha, 3°22'S / 36°41'E, X.–XI.1905, K. Kittenberger ( MNHU); 1 ex., Derema , IX.1891, Conradt S. ( MNHU); 6 ex. , Dodoma, 5°40'S / 35°48'E, XII .2006, F. Kantner (Cka); 1 ex., Mkomazi , 4°38'S / 38°5'E, VIII .1994, G. McGavin, Acacia mellifera canopy sampling ( OUMNH); 1 ex., Mombo—Masinde , 4°53'S / 38°17'E, I.1906, Dr. C. Schröder ( MNHU); 2 ex. , Zanzibar, 6°10'S / 39°11'E, IX.1924, Small ( BMNH); 1 ex. , Kilimandjaro, I.1906, Dr. Chr. Schröder ( MNHU); 1 ex., Magamba Berge, 5°31'S / 38°04'E, I.1905, Dr. Chr. Schröder ( MNHU); 2 ex., Kibonoto , 3°00'S / 37°20'E, VI .1905, Sjöstedt ( MNHU); 1 ex., Gomba , 1912, Inst. Amani ( MNHU); 1 ex. , Kilimandjaro, 5°22'S / 38°02'E, X.1991, C. Schütz (CBe); 1 ex. , Morogoro, 6°49'S / 37°40'E, II.1987, S. Mahunka & A. Ziesi ( HNHM); 1 ex., Mtida, XI.1910, Dr. Holtz ( MNHU); 1 ex., Ukerewe , 2°09'S / 32°52'E, Father Conradt ( NMK) .— Togo. 5 ex., Agou , 6°52'N / 00°43'E, VI .1963, Mme.Y. Schach ( MRAC); 7 ex., Bismarckburg , 8°15'N / 0°55'E, XI.1892 – VI .1893, L. Conradt ( MNHU); 1 ex., Missahoue , 6°59'N / 0°40'E, XI.1893, E. Baumann ( MNHU); 1 ex., Yob, V .1963, Mme. Y. Schach ( MRAC); 2 ex., Togo ( DEIS); 1 ex., Togo , 1892–1893, L. Conradt ( MNHU) .— Uganda. 1 ex., Budongo Forrest , 1°45'N / 31°35'E, I.1997, Th. Wagner ( ZFMK); 2 ex. , Kibale Forrest, I.1984, M. Nummelin ( MZHF); 2 ex., Semliki Forrest, 0°48'N / 30°8'E, II.1997, Th. Wagner ( ZFMK); 1 ex., Kalinzu Forest , X.1948, T . H. E. Jackson ( NMK); 1 ex., Busana , 0°46'N / 32°58'E, III .1925, G. B. R . Hancock ( NMK); 1 ex., Kakumiro, 0°47'N / 31°18'E, I.1923, H. Hargreaves ( NMK); 1 ex., Uganda , I.1922, H. Hargreaves ( NMK) .— Zambia. 1 ex., Kampala, 14°25'S / 31°37'E, A. F. J. Gedye ( NMK); 5 ex., Namwala , 15°45'S / 26°27'E, III .1913, H. C. Dollman ( BMNH) .— Zimbabwe. 2 ex., Mazowe Dam , 17°31'S / 30°59'E, VII .1998, Marlin E. Rice ( TAMU) .