Heteromeringia paraphalloides, Sasakawa, Mitsuhiro, 2011

Sasakawa, Mitsuhiro, 2011, Oriental species of clusiid flies (Diptera: Clusiidae: Clusiinae), Zootaxa 3038, pp. 1-28 : 21-22

Heteromeringia paraphalloides sp. nov.

( Figs 32–35 )

Type material. Holotype male ( BPBM 17250), Vietnam, 15 km NW of Dalat, 1850 m, 5. V. 1960, L. W. Quate. Paratypes: Ƥ, Vietnam, Dilinh (Djiring), 27. IX.–14. X. 1960, C. M. Yoshimoto;Ƥ, Malaysia, Sabah, 19 km N of Kalabakan, forest camp, 21. X. 1962, Y. Hirashima.

Diagnosis. Head black except for face, parafacialia, and gena yellow in male; first antennal flagellomere yellowish, with brown apical area broadened on inner side; dc three; all coxae, fore tibia, and tarsus black, fore tarsomeres more or less flattened; surstylus long, with three spinules apically; cerci united; right paraphallus spiny apically.

Description. MALE. Head black but parafacialia, face, and gena yellow, orbit and ventral margin of frontalia yellowish brown, posterior margin of gena brown; frontalia shining but sparsely pollinose on ventral 1/2; orbit and parafacialia slightly whitish pruinose. Antenna brownish yellow; first flagellomere apically brown, extending broadly on inner side, arista black; palpus black. Thorax black; mesoscutum and scutellum slightly gray-dusted, weakly shining; pleura strongly shining, katepisternum slightly gray-dusted. Wing hyaline, slightly infuscated apically beyond midpoint, fading posteriorly beyond M1; calypter with fringe yellowish; halter yellow. Legs yellow, with all coxae, and fore femur, tibia, and tarsus black. Abdomen shining, dark brown to black; epandrium black. Bristles black.

Frons 1.3 times as wide as eye, parallel-sided on dorsal 1/2 but slightly converging ventrally; third or shortest; parafrontalia with ventral part slightly projecting above eye in profile; oc subequal to poc or third or in length; gena 1/4 as high as eye; vi shorter than third or. First antennal flagellomere shorter than broad, rounded apically, pile on flagellomere and pubescence on arista microscopic.

Thoracic chaetotaxy similar to that of H. melanoprotoma . Wing 3.5 mm long; three costal sections in proportion of 42: 10: 5, r-m beyond middle level of cell dm (15: 10), M1 ratio 4.8, ultimate section of CuA1 about 3/4 length of penultimate section. Fore tarsomeres 1–4 broader than those of other legs, but narrower than distal width of tibia.

Surstylus ( Figs 32, 33 ) long, almost as high as epandrium, bearing three spinules on inner apex; cerci completely united, membranous. Hypandrium ( Fig. 34 ) with five setulae; postgonite chitinized marginally, with three setulae. Phallapodeme 200 µm long; phallus 600 µm long, mesophallus ( Fig. 34 ) twisted at base, hypophallus small, right paraphallus broader than left one and distinctly spiny apically, distiphallus tubular and slightly swollen at tip; ejaculatory apodeme ( Fig. 35 ) 390 µm long, 180 µm broad.

Body length 3.6 mm.

FEMALE. Similar to male, but parafacialia, face, and gena brown, frontalia centrally and face whitish pruinose, vi longer than that of male (as long as or), cell r1 almost entirely brownish, narrowly extending to wing apex and ending just behind R4+5, fore coxa and femur yellow but the latter blackish brown on distal 1/2, fore tarsomeres 1–4 slightly broader than distal width of tibia; wing length 3.2–3.8 mm, body length 3.6–4.3 mm.

Distribution. Vietnam, Malaysia (Sabah).

Remarks. This species is distinguished from its known congeners by the black coxae of all legs, the long surstylus, and the spiny paraphallus.

Etymology. The specific name refers to the characteristic paraphallus.

Figures

FIGURES 25 – 35. Male genitalia of Heteromeringia abatanensis sp. nov. (25 – 27, paratype), H. melanoprotoma sp. nov. (28 – 31, holotype), and H. paraphalloides sp. nov. (32 – 35, holotype). See Figs 1 – 3 for explanation of parts; 31, 35, ejaculatory apodeme, lateral view; 27, HP, hypophallus. Scale = 0.1 mm.