Fistulipora arauzensis, Ernst & Fernández & Fernández-Martínez & Vera, 2012

Ernst, Andrej, Fernández, Luis Pedro, Fernández-Martínez, Esperanza & Vera, Carmen, 2012, Description of a bryozoan fauna from mud mounds of the Lebanza Formation (Lower Devonian) in the Arauz area (Pisuerga-Carrión Province, Cantabrian Zone, NW Spain), Geodiversitas 34 (4), pp. 693-738 : 704-706

Fistulipora arauzensis n. sp. ( Figs 7E, F ; 8 A-E; Appendix)

HOLOTYPE. — SMF 21.165 .

PARATYPES. — SMF 21.166- SMF 21.191.

ETYMOLOGY. — The species name refers to Arauz Sur, the locality where the new species was found.

TYPE LOCALITY. — Arauz Sur (Arroyo section), Province of Palencia, NW-Spain (Cantabrian Mountains).

TYPE HORIZON. — Lebanza Formation, Lower Devonian (Pragian).

DIAGNOSIS. — Encrusting colonies with multilayered secondary overgrowths; autozooecial diaphragms few to absent, thin, planar or concave; autozooecial apertures rounded; lunaria small; vesicles abundant both in endozone and exozone, moderately large, completely isolating autozooecia in 1-2 rows, 6-12 surrounding each autozooecial aperture; maculae small, slightly depressed, consisting of vesicular skeleton, spaced widely on the colony surface.

DESCRIPTION

Encrusting colonies with multilayered secondary overgrowths. Multilayered colonies up to 4.8 mm

thick, separate sheets 0.9-3.5mm thick. Autozooecia growing from laminated basal layer. Autozooecial diaphragms few to absent, thin, planar or concave. Autozooecial apertures rounded. Lunaria small, horseshoe-shaped, directed to the nearest macula, consisting of hyaline material. Vesicles abundant both in endozone and exozone, moderately large, box-like, polygonal in tangential section, having flattened roofs, completely isolating autozooecia in 1-2 rows, 6-12 surrounding each autozooecial aperture; in endozone usually higher than those in exozone. Moderately thick layer of granular skeleton on colony surface often developed.Autozooecial walls thick, granular. Maculae small, slightly depressed, consisting of vesicular skeleton, 0.05-0.06 mm in diameter, spaced widely on the colony surface.

COMPARISON

Fistulipora arauzensis n.sp. is similar to F. compacta Astrova, 1964 from the Lower Devonian (Lochkovian) of Ukraine. The new species differs in having larger vesicles (vesicle diameter 0.07-0.018 vs 0.04-0.12 mm in F. compacta ) and wider spaced autozooecia. Fistulipora arauzensis is also similar to F. trifoliata Schlütter, 1889 from the Middle Devonian of the Eifel but differs from it in having smaller lunaria and larger autozooecial apertures (0.14-0.24 vs 0.12-0.20 mm in F. trifoliata ).

Figures

FIG. 7. — A-D, Altshedata gracilis n. sp.; longitudinal section autozooecia and walls, paratype SMF 21.164; D, longitudinal section showing autozooecia with lunaria and style-like projections, holotype SMF 21.150; E, F, Fistulipora arauzensis n. sp., longitudinal sections of multilayered colonies, paratype SMF 21.170. Scale bars: A, B, D, 1 mm; C, 0.2 mm; E, F, 2 mm.

FIG. 8. — A-E, Fistulipora arauzensis n. sp.; A-B, longitudinal section showing autozooecial and vesicles, paratype SMF 21.170; C, D, tangential section showing autozooecial apertures and vesicles, holotype SMF 21.165; E, longitudinal section showing lunarial deposits in exozone, paratype SMF 21.174;F, G, Eridopora sp. longitudinal section showing autozooecia and large irregular vesicles, SMF 21.192. Scale bars: A, F, G, 1 mm; B, C, E, 0.5 mm; D, 0.2 mm.