Hyocephalus aprugnus Bergroth, 1906

Kment, Petr, Hemala, Vladimír & Malenovský, Igor, 2019, Scanning the Hyocephalidae: details of their external morphology with respect to phylogenetic relationships within Eutrichophora (Hemiptera: Heteroptera) *, Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 59 (2), pp. 423-441 : 437-438

Hyocephalus aprugnus Bergroth, 1906

Hyocephalus aprugnus Bergroth, 1906: 648–649 (original description, illustrations of head in lateral view and wing membrane, distribution). HOLOTYPE: ♀ (macropterous): Australia: South Australia, Yorketown (MZHF; see Note on p. 438).

Hyocephalus aprugnus: BERGROTH (1912) : 343 –344 (description of brachypterous female); GROSSBECK (1912): 360 (list of type specimens, erroneously listing brachypterous female described by BERGROTH 1912) as type); BERGROTH (1913): 146 (catalogue); ŠTYS (1964): 231 –248 (redescription of macropterous female, illustrations of habitus and morphological structures, neotype designation, distribution, parasites); GRANT & ŠTYS (1970): 113 –114 (nomenclature); EYLES (1971): 77 (nomenclature); JORDAN (1972):35: fig. 74 (habitus figure); SCHAEFER (1981): 83 (deposition of non-type material), BRAILOVSKY (2002): 43, 47–50 (illustrations of habitus, genital capsule in posterior view and paramere; distribution; host plant; key to species); CASSIS & GROSS (2002): 134 (catalogue, distribution); YANG (2007): 15, 19: fig. 10, 156 (morphology); YAO et al. (2012):3, 4 (phylogeny); WEIRAUCH et al. (2019): 78 (habitus photo).

Hypocephalus aprugnus (subsequent incorrect spelling): YANG (2007): 19 (morphology).

Material examined. AUSTRALIA: WESTERN AUSTRALIA: 8 miles SW of Mt. Ragged, 22.iii.1968, 1 ♀ (brachypterous), I. F. B. Common & M. S. Upton lgt. ( ANIC); 16 miles N of Collie, 7.iv.1968, 1 ♀ (macropterous), I. F. B. Common & M. S. Upton lgt. ( ANIC); Deepdene, 22.iii.1971, 1 J 3 ♀♀ (brachypterous), E. F. Riek lgt. ( ANIC, 1 J → MMBC, 1 ♀ → NMPC); Deep Dene, Karridale, 25.iii.1964, 1 ♀ (brachypterous), 28.xii.1964, 1 ♀ (brachypterous), L. M. O’Halloran lgt. ( ANIC); Manjimup (Parup Block), pitfall trap, 12.–18.iii.1976, 1 J (brachypterous), J. D. Majer lgt. ( ANIC); Prevelly Pk., 8 km W Margaret R., 33.58°S 114.59°E, 30.x.–1.xi.1984, 1 J 1 ♀ (brachypterous), J. & N. Lawrence lgt. ( ANIC, 1 J → NMPC) [1 ♀ habitus photo, 1 J ESEM]; Sorrento Beach, 14.–15.vi.1967, 2 JJ 2 ♀♀ (brachypterous), B. Humphries lgt. ( ANIC, 1 ♀ → MMBC); Thomas R., 23 km NW by W of of Mt. Arid, 33.515°S 123.00°E, 4.–7.xi.1977, 1 ♀ (brachypterous), M. S. Upton & I. Freehan lgt. ( ANIC) [ ESEM].All P. Kment det.

Distribution. Australia: New South Wales ( BERGROTH 1912), South Australia ( BERGROTH 1906, ŠTYS 1964, BRAI- LOVSKY 2002, CASSIS & GROSS 2002), Western Australia ( BRAILOVSKY 2002, CASSIS & GROSS 2002).

Note. BERGROTH (1906) described H. aprugnus based on a single macropterous female, the holotype, but that specimen became missing since then. ŠTYS (1964: 232) redescribed the species based on a series of macropterous females he discovered in HNHM and designated one of them as neotype. However, the original holotype was later recognised among unidentified Coreidae from MZHF (see GRANT & ŠTYS 1970: 113). GRANT & ŠTYS (1970) applied to the International Commission on Zoological Nomenclature for retention of the neotype in preference to the rediscovered holotype, and placing the names Hyocephalidae , Hyocephalus , and Hyocephalus aprugnus on the Official Lists of Names in Zoology. The neotype designation was accepted in the catalogue of CASSIS & GROSS (2002). However, EYLES (1971) argued against the proposal by GRANT & ŠTYS (1970), finding it pointless. We found no Opinion of the ICZN published in the Bulletin of Zoological Nomenclature regarding this case, and none of the three names is included in the Official Lists ( ICZN 2012; G. Lim, pers. comm. 2019). According to Article 75.8 ( ICZN 1999), the designated neotype is set aside on publication of the discovery of the original name-bearing type which retains its validity.