Carabus Linnaeus, 1758

Azadbakhsh, Saeed & Nozari, Jamasb, 2015, Checklist of the Iranian Ground Beetles (Coleoptera; Carabidae), Zootaxa 4024 (1), pp. 1-108 : 21-26

Genus Carabus Linnaeus, 1758

Subgenus Carabus Linnaeus, 1758

Carabus (Carabus) sculpturatus Ménétriés, 1832 = gilanicus Mandl, 1967

Citations: Guissom (Prov. of Gilan; Mandl, 1967a); Province Mazandaran (Ōsawa et al., 2004).

Subgenus Mimocarabus Gehin, 1876

Carabus (Mimocarabus) elbursensis Breuning, 1946

Citations: North of Iran, Takht-e Suleiman (Prov. of Mazandaran; Breuning, 1946); Hasan Keif Village, 2300 m (Prov. of Mazandaran; Lohaj & Mlejnek, 2007).

Carabus (Mimocarabus) khorasanensis Deuve, 1993

Citations: Ala-Dagh, Bojnurd, 30 km North of Ghuchan (Prov. of Khorasan; Deuve, 1993).

Carabus (Mimocarabus) maurus calosomoides Reitter, 1896 = demetrii Semenov, 1903

Studied material: (ZMTU): Damavand, Gosfand Sara, 2008 (Prov. of Tahran); Tavir, and Firooz Kola 2010 (Prov. Mazandaran).

Citations: ToneKabon "Sefidab" (Prov. of Mazandaran; Semenov, 1903); Without exact locality (Bousquet et al., 2003).

Carabus (Mimocarabus) maurus osculatii Osculati, 1844

Citations: South of Shiraz, env. (Prov. of Fars; Osculati, 1844); Shiraz, Beiza, Sepidan (Prov. of Fars; Mandl, 1967a).

Carabus (Mimocarabus) maurus paphius L. Redtenfoacher, 1843 Citations : Farsistan region (prov. of Fars; Redtenfoacher, 1850).

Carabus (Mimocarabus) roseni gonbadensis Deuve, 1991

Citations: 70 km North east of Gonbad-e-Kavus (Prov. of Golestan; Deuve, 1991).

Carabus (Mimocarabus) roseni hemicalosoma Semenov, 1903 Citations : Bojnurd, env. (Prov. of north Khorasan; Semenov, 1903); Gorgan env (Prov. of Golestan; Semenov, 1903).

Carabus (Mimocarabus) roseni roseni Reitter, 1897

Citations: Without exact locality (Bousquet et al., 2003).

Subgenus Gnathocarabus Deuve, 1991

Carabus (Gnathocarabus) kusnetzovi Semenov, 1903

Citations: Bojnurd, env. (Prov. of north Khorasan; Semenov, 1903); Without exact locality (Bousquet et al., 2003); Bojnurd (Prov. of Khorasan; Lassalle, 2007).

Subgenus Limnocarabus Gehin, 1876

Carabus (Limnocarabus) clathratus stygius Ganglbauer, 1890 Citations : Without exact locality (Bousquet et al., 2003).

Subgenus Tomocarabus Reitter, 1896

Carabus (Tomocarabus) rumelicus kurdistanicus Eidam, 1931 = kurdistanicola Deuve 1990

Citations: Without exact locality (Bousquet et al., 2003).

Carabus (Tomocarabus) rumelicus likninensis Lassalle, 2006

Citations: 4Km east of Likbin, 1700 m (Prov. of West Azarbaijan; Lassalle, 2006; Deuve, 2012).

Subgenus Pachystus Motschulsky, 1866

Carabus (Pachystus) tamsi dashtensis Heinz, 1970

Citations: South of Langerud, 2000 m (Prov. of Gilan; Heinz, 1970); Damash village, 1800 m (Prov. of Gilan; Deuve & Wrase, 2013).

Carabus (Pachystus) tamsi eckerleini Heinz, 1970

Citations: Masuleh, 1800 m (Prov. of Gilan; Heinz, 1970); Assalem and khalkhal, 2200 m (Prov. of Gilan; Deuve & Wrase, 2013).

Carabus (Pachystus) tamsi persicola Deuve & Wrase, 2013

Citations: Esquabad, Neyshabour (Prov. of Khorasan; Deuve & Wrase, 2013).

Carabus (Pachystus) tamsi tamsi Ménétriés, 1832

Citations: Talesh (Prov. of Gilan; Ménétriés, 1832).

Subgenus Oreocarabus Gehin, 1876

Carabus (Oreocarabus) cribratus porrectangulus Gehin, 1885 Citations : Abadeh (Prov. of Fars; Samin et al., 2011).

Carabus (Oreocarabus) cribratus submahouxi Deuve, 2008

Citations: Sahand Mt, 2000-2900 m (Prov. of Ardabil; Deuve, 2008).

Carabus (Oreocarabus) gemellatus Ménétriés, 1832 = ingratus Lapouge, 1907

Citations: Chamroud Village, Elburz Mts, R., 1800-2800 m; Forest of Tunekabon, 1250 m (Prov. of Mazandaran; Lapouge, 1907); Without exact locality (Bousquet et al., 2003).

Subgenus Sphodristocarabus Gehin, 1885

Carabus (Sphodristocarabus) adamsi Adams, 1817

= serriae Deuve, 2002

Citations: North of Iran ( Adams, 1817); Kaleybar env. (Prov. of Mazandaran; Deuve, 2002).

Carabus (Sphodristocarabus) bohemani bohemani Ménétriés, 1832

Citations: Azerbaydzhan/ Iran borders region, N-Iran (Provinces Gilan and Mazandaran; Ménétriés, 1832).

Carabus (Sphodristocarabus) bohemani esfandiarii Heinz, 1973

Citations: Sardab-rud, Rudbarak, South of Chalus and Kelardasht (Prov. of Mazandaran; Heinz, 1973).

Carabus (Sphodristocarabus) bohemani esperanzae Heinz, 1970 Citations : Masuleh, Asalam (Prov. of Gilan; Heinz, 1970).

Carabus (Sphodristocarabus) bohemani morvani Morvan, 1976

Citations: Elburz Mt.R., Chorteh, 1800 m, South west of Ramsar (Prov. of Mazandaran; Morvan, 1976).

Carabus (Sphodristocarabus) bohemani rosicolor Deuve, 2001 Citations : Larkhani Mts., 1400 m (Prov. of Gilan; Deuve, 2001).

Carabus (Sphodristocarabus) elegantulus Motschulsky, 1850

= astrabadensis Semenov, 1903

dinae Rapuzzi, 2005

parthicus Lapouge, 1924

sarafrazii Deuve, 2000

Citations: Elburz Mt.R (Motschulsky, 1850); Mts. of Astrabad (Prov. of Golestan; Semenov, 1903); Bojnurd env. (Prov. of Khorasan; Lapouge, 1924); Sari, Kiyasar, Vesmine Mts., 1500-1600 m (Prov. of Mazandaran; Deuve, 2000); Ziarat, 1160 m (Prov. of Golestan; Deuve, 2000); National Park of Golestan, Minu Dasht; Dowlat-abad env, 700 m (Prov. of Golestan; Deuve, 2000); South of Gaz, 36:39N/53:58E (Prov. of Golestan; Rapuzzi, 2005).

Carabus (Sphodristocarabus) hollbergi Mannerheim, 1827 Citations : Province Mazandaran ( Chaudoir, 1842).

Carabus (Sphodristocarabus) separatus bararii Deuve, 2000

Citations: Ramsar env., Ney Dasht, 300-350 m (Prov. of Mazandaran; Deuve, 2000).

Carabus (Sphodristocarabus) separatus haerii Heinz, 1973

= larkhaniensis Deuve, 2001

Citations: East of Rudbar, Dare-dasht (Prov. of Gilan; Heinz, 1973); Gilan env., Larkhani Mts., 1400 m; betw. Siyahkal/Espeyli and Balarud env. (Prov. of Gilan; Deuve, 2001).

Carabus (Sphodristocarabus) separatus korgei Heinz, 1970 = delirensis Deuve, 2000

Citations: south west of Chalus, Delir, 36˚21’N/51˚08’E (Prov. of Mazandaran; Deuve, 2000); South of Ramsar (Prov. of Mazandaran; Lassalle, 2001).

Carabus (Sphodristocarabus) separatus persa Lapouge, 1924 = sahelae Lassalle, 2001

longicollis Mandl, 1955

Studied material: (ZMTU): Nowshaher, Khirood forest (Prov. of Mazandaran).

Citations: Elburz Mt. R., Ramsar env,. to Sari env., Kelardasht (Prov. of Mazandaran; Lapouge, 1924); Elburz Mts. R ( Mandl, 1955); Galugah, 1800 m (Prov. of Mazandaran; Lassalle, 2001); Hasan Keif village, 2300 m (Prov. of Mazandaran; Lohaj & Mlejnek, 2007).

Carabus (Sphodristocarabus) separatus samaorum Rapuzzi, 2005

Citations: 20 km South of Gorgan, Nahar Horan (Prov. of Golestan; Rapuzzi, 2005).

Carabus (Sphodristocarabus) separatus separatus Lapouge, 1907 Citations : Talesh, 2500 m (Prov. of Gilan; Lapouge, 1907); Asalom, Masuleh "Talysh" (Prov. of Gilan; Mandl, 1967b).

Carabus (Sphodristocarabus) separatus tarkhaniensis Deuve, 2001 Citations : Larkhani, 1400 m (Prov. of Gilan; Deuve, 2000).

Subgenus Megodontus Solier, 1848

Carabus (Megodontus) persianus domradensis Deuve, 2008

Citations: Javaher Deh valley, Domrad Mts., 1100 m (Prov. of Mazandaran; Deuve, 2008).

Carabus (Megodontus) persianus espeyliensis Deuve, 2001

Citations: W-Elburz Mt. R, betw. Siyakhal:Balarud Mts (Prov. of Gilan; Deuve, 2001).

Carabus (Megodontus) persianus javahardehensis Deuve, 2000

Citations: 20 km South west of Ramsar, Javahar Deh (Prov. of Mazandaran; Deuve, 2000).

Carabus (Megodontus) persianus neydashtensis Deuve, 2000

Citations: South of Ramsar, Neydasht, 800 m (Prov. of Gilan; Deuve, 2000).

Carabus (Megodontus) persianus persianus Roeschke, 1896 = damashensis Deuve, 2006

Citations: Talesh Mts, btw Iran-Azarbaijan (Prov. of Gilan; Roeschke, 1896); Damash, East of Rudbar (Prov. of Gilan; Deuve, 2006).

Carabus (Megodontus) stroganowi ebrahimii Deuve, 2000

Citations: South of Nur, Ruyan, Kodirsar, 1200 m (Prov. of Mazandaran; Deuve, 2000).

Carabus (Megodontus) stroganowi lavidzhensis Deuve, 2001

Citations: 25km South east of Nur, Lavidzh (Prov. of Mazandaran; Deuve, 2001).

Carabus (Megodontus) stroganowi luisae Rapuzzi, 2005 Citations : Province Gilan ( Rapuzzi, 2005).

Carabus (Megodontus) stroganowi mofidii Deuve, 2000

Citations: Kiyasar env., 1600 m (Prov. of Mazandaran; Deuve, 2000).

Carabus (Megodontus) stroganowi morgani Lapouge, 1907

= heinrichi Mandl, 1966

Studied material: (ZMTU): Kolijak, 2012 (Prov. of Mazandaran).

Citations: Elburz Mts. R, TuneKabon, Khanian, 1250 m (Prov. of Mazandaran; Lapouge, 1907); Psh-Kuh, 400- 1800 m (Prov. of Lurestan; Mandl, 1966 sub heinrichi); Without exact locality (Bousquet et al., 2003).

Carabus (Megodontus) stroganowi pseudopersianus Deuve, 2001 Citations : Rudbar env. (Prov. of Gilan; Deuve, 2001).

Carabus (Megodontus) stroganowi stroganowi Zoubkoff, 1837 = galugahensis Lassalle, 2001

Citations: Astrabad (Prov. of Golestan; Zoubkoff, 1837); btw, Behshahr/Gorgan, Mts, of Galugah, 1800 m (Prov. of Mazandaran; Lassalle, 2001).

Carabus (Megodontus) stroganowi tiedemanni Breuning, 1935

Citations: Asalem and Masuleh (Prov. of Gilan; Breuning, 1934).

Carabus (Megodontus) stroganowi transfuga Semenov, 1898 = transcurrens Csiki, 1927

Citations: North of Iran, Talesh Mt.s (Prov. of Gilan; Semenov, 1898).

Subgenus Procerus Dejean, 1828

Carabus (Procerus) scabrosus elbursianus Mandl, 1958 Citations : Elburz Mts. R, Tacht-e Suleiman, Sardab-Tal "Hasankif", 10-1400 m (Prov. of Mazandaran; Mandl, 1958).

Carabus (Procerus) scabrosus farnazae Deuve, 2006 Citations : Without exact locality ( Deuve, 2006).

Carabus (Procerus) scabrosus sommeri Mannerheim, 1844 Citations : Province Mazandaran ( Chaudoir, 1844).

Subgenus Procrustes Bonelli, 1810

Carabus (Procrustes) clypeatus clypeatus Adams, 1817 Citations : North of Iran ( Adams, 1817).

Carabus (Procrustes) clypeatus kurnakovi Kryzhanovskij, 1968 Citations : Province Mazandaran ( Chaudoir, 1842).

Carabus (Procrustes) talyschensis luctuosior Deuve, 1994

= luctuosus Zoubkoff, 1837

Citations: Astarabad (Prov. of Golestan; Zoubkoff, 1837); Babol (Prov. of Mazandaran; Mandl, 1953 sub luctuosus ); Gorgan (Prov. of Golestan; Mandl, 1967a sub luctuosus ).

Carabus (Procrustes) talyschensis talyschensis Ménétriés, 1832

Studied material: (ZMTU): Kolyak, 1626m. 2008 (Prov. of Mazandaran).

Citations: Talesh (Prov. of Gilan; Ménétriés, 1832); Gorgan env. (Prov. of Golestan; Afshari & Khormali, 2014).

Subgenus Lamprostus Motschulsky, 1866

Carabus (Lamprostus) calleyi nigrinus Motschulsky, 1866

Citations: 30 km East of Ahar 1500 m; North of Kaleibar, 1750 m; Davehsar-dagh North of Ahar 1700 –2300 m; Kholoma Yurm North of Ahar 1700 m; Varid East of Ardebil 1500–1800 m; Sabalan, 2000 m, 2600–3200 m; Talysch, 1600 m (Prov. of East Azarbaijan; Heinz, 2002); Btw. Astara/Ardebil (Prov. of Gilan; Heinz, 2002).

Carabus (Lamprostus) calleyi prasinus Ménétriés, 1832

Studied material: (ZMTU): Lenkoran, 2012 (Prov. of Azarbaijan).

Citations: Talesh (prov. of Gilan; Ménétriés, 1832); Rostamabad 1700 – 1900 m (Prov. of Gilan; Heinz, 2002).

Carabus (Lamprostus) calleyi rechingerae Mandl, 1972

Citations: Dasht-Moghan (Prov. of East Azarbaijan; Mandl, 1972b); Dasht-e Moghan (Prov. of Ardabil; Deuve, 2001); Parsabad (Prov. of West Azarbaijan; Deuve, 2001).

Carabus (Lamprostus) cylindricus Lapouge, 1914

= taveni Heinz, 2002

cylindricior Deuve, 1994

Citations: Elburz Mts. R (Deuve, 2001); Dailiman 1600–1800 m (Siah-Sar) and Pass Masuleh/Ali-abad 2000 m; Pass Ardebil/Astara 1500 m (Prov. of Gilan; Heinz, 2002); Pass Khalkhal/Assalam, S. Seite, 2200 m; Sabalan 2600–3200 m (Prov. of East Azarbaijan; Heinz, 2002).

Carabus (Lamprostus) inseriatus Mandl, 1955

Citations: West of Ardabil, Nir env (Prov. of Ardabil; Mandl, 1955); West of Ardabil, 1900 m (Prov. of Ardabil; Deuve, 2001); Nir West of Ardebil 1900 m; Davehsar dagh North of Ahar 1700–2300 m (Prov. of East Azarbaijan; Heinz, 2002).

Carabus (Lamprostus) guycolasianus colorescens Deuve, 2001 Citations : Elburz Mts. R (Deuve, 2001).

Carabus (Lamprostus) guycolasianus guycolasianus Deuve, 2001 Citations : 100 km South of Chalus, 2300 m (Prov. of Mazandaran; Deuve, 2001); Rostamabad (Prov. of Gilan; Deuve, 2001).

Carabus (Lamprostus) ledouxi Deuve, 2001

Citations: Hero-Abad Mt, Charasu Dagh, 2000-2300 m (Prov. of East Azarbaijan; Deuve, 2001); Asbhoni village, 2000 m; Talych, Elburz Mts. R; Astara, 1900 m (Prov. of Gilan; Deuve, 2001); Ardabil (Prov. of Ardabil; Deuve, 2001); West of Ahar, 1400 m; Saqqez (Prov. of East Azarbaijan; Deuve, 2001).

Carabus (Lamprostus) seroulikbin Cavazzuti, 2008

Citations: 4 km east of Likbin, 1660-1700 m and 12 km south of Derou 1950-2000 m (Prov. of West Azerbaigan; Cavazzuti, 2008).