Peltonotus sisyrus Jameson and Wada, 2004

Jameson, Mary Liz & Wada, Kaoru, 2004, Revision of the genus Peltonotus Burmeister (Coleoptera: Scarabaeidae: Dynastinae) from Southeastern Asia, Zootaxa 502 (1), pp. 1-66 : 44-46

Peltonotus sisyrus Jameson and Wada , n. sp.

( Figs. 28 , 49a–c , 62 , 65 )

Type Material. Holotype male housed at FUJI with following label data and male genitalia and mouthparts mounted beneath specimen: a) "near Aceh N. SUMATRA 1993. NOV." (type face), b) " Fujioka­Collection Peltonotus­G­1" (typeface and handwritten, yellow label), c) our holotype label . Allotype female housed at FUJI with following label data: a) "near Brastagi NORTH SUMATRA JUN. 1995." (type face), b) "Fujioka­Collection Peltonotus­G­2" (typeface and handwritten, yellow label), c) our allotype label .

Description. Holotype male. Length 16.1 mm. Widest width 7.7 mm. Color: Head, pronotum, scutellum, pygidium, elytron, and venter castaneous. Elytra with weak iridescent bloom Head: Surface of frons with base impunctate (middle) to sparsely punctate (laterally), middle frons to apex moderately densely punctate, more dense laterally; punctures simple, multisetigerous; setae minute (1–12+ per puncture) and moderately long adjacent to eye (0–1 per puncture). Surface of clypeus moderately densely punctate, more dense laterally; punctures simple, multisetigerous; setae minute (1–12+ per puncture) and short near margin (0–1 per puncture). Clypeus laterally weakly bowed, apex truncate, corners square, beaded; bead not weakly arcuate posteriorly. Labrum broadly narrowly emarginate at middle. Mandible with external edge rounded, inner apex with 2 teeth. Mentum with apical half triangular, notched at middle; palpomere 2 dorsoventrally flattened, about 3 times width of palpomere 1, setose; setae dense, moderately long, rufous, weakly thickened, not curled. Maxilla ( Fig. 28 ): mala with dense lamellate setal brush; stipes with setae dense, long, flattened at apex, not curled at apex; palpomere 2 with weak internomedial bump. Antennal club subequal to segments 2–7. Pronotum: Bead lacking anterior to scutellum. Surface moderately densely punctate; punctures simple, multisetigerous laterally; setae minute (1–7 per puncture) and short (0–1 per puncture). Lateral margin lacking long setae. Elytral sutural length: About 3.5 times length of scutellum. Elytra: Surface with 7 poorly developed, punctate, longitudinal striae between suture and humerus; punctures ocellate, moderate in size, moderately dense, multisetigerous apically and laterally; setae minute (1–20+ per puncture). Intervals similarly sculptured. Propygidium: Surface shagreened and moderately densely punctate; punctures simple, unisetigerous (short at base to mid­disc) and multisetigerous (minute at apex). Pygidium: Surface densely punctate; punctures ocellate, multisetigerous; setae minute (7–20+ per puncture) and long at apex (1–2 per puncture). Venter: Prosternal keel elongate; apex projecting anteriorly at about 90° with respect to ventral plane, produced to about 3/4 of protrochanter, truncate. Legs: Foretibia of male bidentate; lateral margin with short, dense setae. Foretarsomere 5 subequal in length to tarsomeres 1–4, greatly thickened; foretarsomeres 2–4 with apices expanded (moreso in 3), dorsal and ventral apices of tarsomeres 1–4 clothed with small patch of dense, short setae. Foreclaws with inner claw broadly curved, about 3 times thicker than outer claw; outer claw elongate­arcuate, about 1/2 the length of inner claw; empodium bulbous at base. Meso­ and metatibial claws of male with 2 setae, claw angled toward venter, about 1/2 length of metatarsomere 5. Metatibia of male with apical spurs more or less straight; ventral spur produced to middle of metatarsomere 1, dorsal spur produced to apex of metatarsomere 2. Parameres: Fig. 49a–c .

Allotype Female. Differs from the holotype in the following respects. Length 16.4 mm. Widest width 7.6 mm. Color: Head, pronotum, scutellum, propygidium, pygidium, and venter black; elytra black with iridescent bloom. Head: Surface of frons densely punctate, multisetigerous; setae minute (1–12+ per puncture) and short (0–1 per puncture). Surface of clypeus densely punctate, multisetigerous; setae minute (1–12+ per puncture) and short (0–1 per puncture). Pronotum: Punctures unisetigerous apicolaterally; short (0– 1 per puncture). Propygidium: Punctures unisetigerous with line of short setae near apex. Elytron: Epipleuron of female in ventral view expanded, broadly incised at sternite 4; in dorsal view expansion moderately developed ( Fig. 62 ). Legs: Foreclaws subequal 1/2 length of foretarsomere 5, claw angled toward venter.

Diagnosis. Peltonotus sisyrus is one of only two Peltonotus species that possess a mentum that is triangular in the apical half ( Fig. 28 ; shared with P. deltamentum ). It is easily separated from other species of Peltonotus based on the greatly enlarged fifth foretarsomere in the male and its distribution in northern Sumatra. In addition, the surface of frons and clypeus is multisetigerous; labrum broadly narrowly emarginate at middle; mandible with external edge rounded; male with foretarsomere 5 greatly thickened and apices of tarsomeres 1–4 clothed with small patches of dense, short setae; outer foreclaw elongate­arcuate; and epipleuron of female in ventral view expanded, broadly incised at sternite 4 ( Fig. 62 ).

Etymology. The specific epithet “ sisyrus ” is Greek for “shaggy garment” and refers to the dense short and long setae on the head of this species.

Distribution ( Fig. 65 ). Sumatra, Indonesia.

Locality records (2 specimens) from FUJI .

INDONESIA (SUMATRA). Aceh Province (2): Banda Aceh, Brastagi .

Temporal Data. June (1), November (1).

Figures

FIGURE 65. Distribution of Peltonotus species in Southeast Asia and Sundaland.

FIGURES 19–28. Mentum, dorsal view of: 19) P. deltamentum, 20) P. gracilipodus, 21) P. kyojinus, 22) P. malayensis, 23) P. podocrassus, 24) P. nasutus, 25) P. nethis, 26) P. pruinosus, 27) P. similis, 28) P. sisyrus.

FIGURES 49–50. Male genitalia in dorsal (parameres only [a]) and lateral views (parameres and phallobase [b–c]) of: 49) P. sisyrus, 50) P. vittatus.

FIGURES 61–64. Elytral epipleuron (gray, ventral view) with position of metacoxa of: 61) P. similis, 62) P. sisyrus, 63) P. suehirogarus, 64a–b) P. vittatus.