Nepalomyia singaporenis, Grootaert, 2013

Grootaert, Patrick, 2013, New Nepalomyia Hollis (Diptera: Dolichopodidae) From Singapore And South Malaysia, Raffles Bulletin of Zoology 61 (1), pp. 103-116 : 112-113

Nepalomyia singaporenis , new species

( Figs. 21–24 , 25–27 )

Material examined. — Holotype male, SINGAPORE: Sime Forest , 1 Apr.2005, forest, Mal 1, (MS name: Si 899; reg. 25040, Si 899, coll. P. Grootaert).

Paratypes: 1 male, Nee Soon , 8 Jul.2005, swamp forest, Mal 2, (reg. 25206, Si 894, coll. PG) ; 2 males, Sime Forest , 27 Apr.2005, forest, Mal 1, (reg. 25093, Si 902, coll. PG), male in genbank ; 1 male, Sime Forest , 3 Jun.2005, forest, Mal 1, (reg. 25134, Si 906, coll. PG) ; 1 male, Sime Forest , 17 Jun.2005, forest, Mal 1, (reg. 25136, Si 914, coll. PG) ; 1 male, Sime Forest , 13 May 2005, forest, Mal 2, (reg. 25130, Si 943, coll. PG) ; 5 males, Sime Forest , 6 May 2005, forest, Mal 2, (reg. 25128, Si 1063, coll. PG) ; 1 male, Sime Forest , 3- Jun. 2005, forest, Mal 2, (reg. 25135, coll. PG) ; 1 male, Sime Forest , 3 Dec.2005, forest, Mal 1, (reg. 25434, coll. PG) .

In genbank as Nepalomyia Si 899 ( Lim et al., 2009).

Etymology. — The name refers to the type locality of the species.

Diagnosis. — Small species with triangular third antennal segment almost 3 times as long as wide (2.7×); arista 1.6 times as long as third segment. Basal fifth of vein R2+3 thickened. Hind metatarsus with a dorsal ridge.

Male: Body length: 2.1 mm; wing length: 1.95 mm. Head. Vertex brownish black, surpassing level of upper eye margin. Occiput concave. Face shining metallic blue, clypeus grey dusted. Face above about twice as wide as second antennal segment, narrowing downwards. Clypeus very feebly bulging out. Palpus dark brown, with a black apical bristle. Rostrum yellowish brown. Ocellar tubercle prominent. 2 strong black ocellars, 2 postverticals, nearly as long as the postocellars, but less strong; 2 tiny postocellars. Postocular bristles uniseriate, black. Antenna brown: Second segment with a circle of short marginal bristles and with 1 or 2 long dorsal bristles. Third segment elongate triangular ( Fig. 21 ), 2.7 times as long as wide. Arista 1.6 times as long as third segment, base pale brownish, rest almost white.

Thorax brown, not shining, pleurae hardly lighter coloured. A shallow prescutellar flattened area. Postscutellum with a median mound. Acr biseriate, long, half as long as dc. 5 dc (growing longer posteriad). 1 long humeral and 1 only slightly shorter posthumeral, 1 short presutural, 2 long postsutural; 1 very long pre-alar; 1 very long post alar, 2 long notopleurals; scutellum with 2 strong marginals with a short hair at side. 1 very long black propleural bristle, just above coxa.

Legs yellow with mid and hind coxae brown. Bristles and hairs black. Fore coxa anteriorly with short hairs ending in 4 long bristles near tip of coxa. Fore femur without bristles, bearing only 2 very short, hairlike preapical pv. Fore tibia with short apical bristles. Mid coxa with numerous anterior bristles and 1 very long black bristle below middle. Mid femur with a strong anterior preapical, no posterior preapical. Mid tibia: 2 strong ad, 2 weaker pd; 3 strong bristles in apical crown. Mid tarsomeres with a pair of black apical bristles. Hind coxa with a black exterior bristle, half as long as the one on mid coxa. Hind femur with an anterior preapical; no posterior preapical and no other bristles. Hind tibia ( Fig. 22 ) with ad on basal quarter and distinct dorsal just beyond middle followed by a few less distinct dorsal, a strong anterodorsal preapical. No real apical crown of bristles, only a distinct ventral. Posterodorsal comb short ( Figs. 23, 24 ), with a black denticle below. First segment of hind tarsus a little longer than half as long as second segment, with a elongate ridge (“spur”) at base. All hind tarsomeres with black apical bristles. All claws minute, symmetrical.

Wing faintly brownish tinged with brown veins. Basal fifth of vein R 2+3 thickened. Joint of R 2+3 and R 4+5 a little swollen. Apical part of M 3+4 about 2.5 times as long as tp. Anal vein pale brownish near base, but fading and not reaching wing border. Squama brown with fan of long, black cilia. Halter with pale knob and pale stalk.

Abdomen brown. Hairs and bristles on terga short, all marginals distinct, slightly longer as tergites are wide. Sterna pale brownish with long marginals. Tip of surstyli and aedeagus hidden in sternites 4 and 5. Genital capsule ( Fig. 25 ) globular, dark brown. Cerci elongate, with a strong, yellowish bristle near tip. Surstyli elongate, brown in contrast to paler cerci; about half as long as genital capsule is long. Hypandrium ( Fig. 26 ) yellowish; tip with 4 long narrow teeth. Aedeagus ( Fig. 27 ) yellowish. A long pale, epandrial bristle present, inserted on a short tubercle; with a short bristle at base.

Female: unknown.

Remarks. — N. singaporenis , new species seems to be related to N. horvati Wang & Yang (2004) in that both species have a long triangular third antennal segment, 2.7 times as long as wide in N. singaporenis , but 4.6 times in N. horvati . Both species have the basal spur on hind metatarsus rim-like. The basal portion of vein R2+3 is not swollen in N. horvati as it is in N. singaporensis , new species The genitalia are similar in general structure but much different in the details. Cercus in N. singaporensis bears a broad leaf-like bristle. This bristle is slender and much longer in N. horvati .

Wang & Yang (2004) assigned N. horvati to the N. longa - group, and due to the many similarities, N. singaporensis , new species should also belong to this group. However N. singaporensis , new species does not have basal tubercles on the cercus or at least it is not recognised as such here in this study.

Figures

Figs. 21–24. Nepalomyia singaporensis, new species: 21, antenna; 22, hind tibia and basitarsus; 23, hind basitarsus dorsally; 24, hind metatarsus laterally.

Figs. 25–27. Nepalomyia singaporensis, new species: 25, postabdomen; 26, tip hypandrium; 27, tip aedeagus.