Ilex wiesaensis MAI

Dedicated in memory of the late FrantiŠek Holý (1935 - 1984), an eminent Czech palaeobotanist, Holý, František, Kvaček, Zlatko & Teodoridis, Vasilis, 2012, A Review Of The Early Miocene Mastixioid Flora Of The Kristina Mine At Hrádek Nad Nisou In North Bohemia (The Czech Republic), Acta Musei Nationalis Pragae Series B 68 (3 - 4), pp. 53-118 : 84-85

Ilex wiesaensis MAI

Pl. 9, fig. 19

1970b Ilex wiesaensis MAI , p. 458, pl. 60, figs 14-18 (Wiesa).

Stone asymmetrically ellipsoidal, 4.3 mm long, 1.6 mm wide, ventral edge straight, dorsal side slightly convex, with 6 very thin ribs running throughout the whole side and rarely anastomosing, lateral sides with 2 or more distinct ribs ( Holý 1975, p. 67, pl. 12,fig.11).

D i s c u s s i o n: According to Holý (1975) similar endocarps are produced by I. ambigua CHAPM. living in North America.

M a t e r i a l: One stone, G 3061.

Adoxaceae TRAUTV.

Sambucus L. Sambucus pulchella C. et E.M. REID

Pl. 9, figs 20-21

1915 Sambucus pulchella C. et E.M. REID, p. 135, pl. 17, figs 7-10 (Reuver).

1977a Sambucus pulchella C. et E.M. REID; Holý, p. 113 (Hrádek/N., Kristina Mine).

Seeds anatropic, elongate ovoid, 1.9–2.4 mm long, 1.1–1.5 mm wide, apex slightly acute, with a tiny hilum shifted sideward, base rounded, surface granulate due to transverse densely spaced rows ( Holý 1975, p. 94, pl. 20, fig. 10).

D i s c u s s i o n: According to Holý (1975) seeds of Sambucus parvula and S. colwellensis ( Chandler 1963b, Upper Headon) are smaller, with more distinct transversal striae, those of S. muddensis ( Chandler 1963a, Mudeford) are similar in size, but the surface is coarsely rugulate with deep furrows. The material from Hrádek best matches the records from the Pliocene of Reuver, Swalmen and Brunssum. Similar populations were described from Krościenko ( Szafer 1947, as S. cf. ebulus L.) and western Siberia ( Dorofeev 1963a), the latter differing in its coarse sculpture ( Holý 1975). According to Czaja (2003) similar seeds to this species were later published from Köflach ( Meller 1998, as Sambucus sp. ) and Berzdorf matching those produced by S. nigra L. and S. racemosa L. native in Europe ( Czaja 2003).

M a t e r i a l: 4 seeds, G 3032, 8855-56.

Angiosperms fam. inc.? Monocotyledonae gen. et sp. indet.

Pl. 14, fig. 8

Leaf (?) fragment with parallel venation. Epidermis on both sides thickly cutinized, finely granulate and in thicker zones striate, reflecting trigonal to polygonal cell outlines 15–35 µm in diameter, partly parallel, straight-walled, some cells thickened, stomata widely scattered, perpendicularly arranged to cell length, amphicyclic, guard cell pairs thinly cutinized, 30 µm long and 40 µm wide, with thickened ledges bordering boat-shaped pore, subsidiary cells narrow, half-moon shaped, in 2–3 circles per 8 cells, thickly cutinized. Strong trichome bases simple, 20–25 µm in diameter, solitary, irregularly disposed.

D i s c u s s i o n: Dispersed cuticles with similar structure were described as Freycinetia rhenana WEYLAND (1957) from Rhineland and Monocotylophyllum lusaticum JUCHNIEWICZ (1975) from Turów showing less cutinized stomata and less complicated stomatal complexes. According to Kvaček and Wilde (2006) the tissue fragment in the former case belongs to the stalk of a dicot leaf. The affinity of the above described structure remains dubious.

M a t e r i a l: Dispersed cuticle, G 9398 (KR 324).

Symplociphyllum KVAČEK et BŮŽEK Symplociphyllum weylandii KVAČEK et BŮŽEK

Pl. 4, figs 1-4, pl. 14, figs 9-10

1959

1966

Illicium lusaticum (JÄHNICHEN) KRÄUSEL et WEYALND sensu KRÄUSEL et WEYLAND , p. 106, pl. 20, figs 12-15, text-fig. 6-7 (non Kadsura lusatica JÄHNICHEN nec Illicium lusaticum (JÄHNICHEN) KRÄUSEL et WEYLAND pro nomen) (Düren).

Symplociphyllum weylandii KVAČEK et BŮŽEK, p. 293, pl. 3, figs 4-5, pl. 4, fig. 3 (Hrádek/N., Kristina Mine).

Incomplete leaf, probably elliptic to ovate, 35 mm long, 17 mm wide, apex and base not preserved, margin entire, venation brochidodromous, midrib strong and straight, secondary veins very thin, straight, alternate to subopposite, originating at an angle of 40–60°, looping at the margin, tertiary veins perpendicular, straight to sinuous, venation of higher orders poorly preserved. Adaxial epidermis medium cutinized, smooth, non-modified cells polygonal, irregularly disposed, anticlinal walls coarsely wavy and sinuses with indistinct lens-shaped tickenings, scattered solitary simple rounded trichome bases, abaxial cuticle thinly cutinized, almost smooth, only faintly radially striate around some stomata, non-modified cells of the same form and size as adaxially, but without thickenings in sinuses, stomata incompletely cyclocytic, subsidiary cells 3–4 in one circle, slightly darker, guard cell pairs broadly oval to circular, 25–27 µm long and 20–25 µm wide, stomatal ledges short and not thickened, bordering oval pore.

D i s c u s s i o n: The stomatal type in the original diagnosis ( Kvaček and Bůžek 1966) was wrongly interpreted as paracytic. The same type of dispersed cuticles with stomata was described as Coronicutis hartauensis ROSELT et SCHNEIDER (1969) from the nearby locality of Hartau in Germany (abaxial leaf side only) and Myrsine miocenica JUCHNIEWICZ (1975) from Turów. So far the affinity has not been resolved. The Symplocaceae family is ruled out by the stomatal type. The affinity to Myrsinaceae R. BR. requires more detailed comparisons. A fragment with similar cuticle structure from the mastixioid flora of the Oberdorf Mine (Kovar-Eder et al. 2001) differing mainly in the striate adaxial cuticle was assigned to Ternstroemites ( T. waltheri KOVAR-EDER ).

M a t e r i a l: Leaf compression on slide, G 9399 (KR 182).