Chaetopodella latitarsis, Hayashi & Papp, 2007

Hayashi, T. & Papp, L., 2007, Oriental Species Of Chaetopodella Duda (Diptera: Sphaeroceridae), Acta Zoologica Academiae Scientiarum Hungaricae 53 (2), pp. 117-130 : 120-122

Chaetopodella latitarsis sp. n.

( Figs 8–12 )

Holotype male ( NIID): VIET NAM, Mt. Phang Si Pang , N. rdg., 1,950–1,970 m, Lai Chau Prov., Oct. 4–6, 1999, T. Hayashi – Cattle dung.

Paratypes VIET NAM: 36 males, 16 females, same data as holotype ( NIID, 2 males, 2 females HNHM) ; 33 males, 19 females, Mt. Phang Si Pang , N. rdg., 1,950 –1,970 m, Lai Chau Prov., Oct. 3–4, 1995, H. Kurahashi ( NIID) ; 1 female, De O Quy Ho, 1,750 m, Sa Pa , Lao Cai Prov., June 3, 1999, H. Kurahashi ( NIID) ; 10 males, 3 females, ditto, Oct. 5, 1999, T. Hayashi ( NIID) ; 1 male, 2 females, Sa Pa , 1,500 m, Lao Cai Prov., Oct. 2, 1995, H. Kurahashi ( NIID) ; 2 males, 1 female, Sin Chai , “Legendary place”GEOCODE??, Nov. 15, 2003, No. 6, along forest path & creek, leg. Földvári, Peregovits, Kőrösi ( HNHM, 1 male gen. prep.) . THAILAND: 1 male, 2 females, Chiang Dao , 1,200 m, Oct. 25, 2001, T. Hayashi ( NIID) ; 1 female, Doi Angkhang (nr. Fang ), 1,700 m, Oct. 27, 2001, T. Hayashi ( NIID) .

Measurements in mm: body length 2.20 mm (holotype), 2.09–2.25 mm (paratype males),

2.09–2.58 (paratype females), wing length 2.30 mm (holotype), 2.20–2.35 (paratype males), 2.31–2.51 (paratype females), wing width 0.92 (holotype), 0.88–0.97 (paratype males), 0.96–1.01 (paratype females).

Head: Mostly blackish brown, heavily microtrichose; silvery spots around bases of macrochaetae; anterior part of frons reddish brown; ocellar triangle silvery; gena pale yellow in male and reddish brown in female, face pale yellow in male and brown to dark brown in female; gena with a batch of dark minute setae directed downwards near posterior end, they are especially prominent in male; 3 weak ifr; anterior ors about as long as 1/2 length of posterior ors; eye oval, its longest diameter about 3.0 times as long as smallest genal width; antenna dark brown, base of 1st flagellomere reddish brown; arista long, about 4.2 times as long as antenna, shortly ciliate.

phallapodeme laterally, 12 = 5th sternite, ventral view. Scale: 0.1 mm for all

Thorax: Mostly blackish brown; mesonotum with 3 broad velvety stripes on dorsocentral lines and acrostichal area, other part heavily greyish microtrichose, 6 rows of ac microsetae in front of suture, 4 dc, posterior 3 very strong; anepisternum, katepisternum, anepimeron and meron dark brown, strongly microtrichose; pale yellowish line on notopleura; 2 kepst, both weak; scutellum velvety black, somewhat wider than long; 2 sc long, apical one about 1.8 and lateral one about 1.0 times as long as scutellum.

Wing: Yellowish, veins yellowish brown; C somewhat darker, not extended beyond apex of R 4+5, first sector with a little stronger setae than those of second and third sectors; R 4+5 gently bent up to C; C-index (Cs2: Cs3) = 1.55–1.85; R-M – dM-Cu: dM-Cu = about 2.5; alula broad and rounded; halter yellowish brown to reddish brown.

Legs: Yellowish brown to dark brown (generally darker in female). Male fore tibia somewhat swollen, with anterior incision on apical 1/3 as in that of Spelobia clunipes (MEIGEN) , fore tarsus dark brown and flattened, each tarsomere with long dense lateral setae. Fore femur with a dorsal seta on basal 2/3; mid femur with a ventral row of 5 setae on basal 1/ 5 in male; mid tibia with 3 dorsal setae on basal 1/4,1/3 and 3/4, 1 rather strong anterodorsal seta on 2/3, 2–3 weak posterodorsal setae on 1/3–1/2, rather strong posterodorsal seta on 2/3, 1 strong apicoventral seta, no ventral seta in male, and a strong ventral seta on basal 1/ 2 in female.

Abdomen: Preabdominal sternites broad, sternite 4 quadrate (0.30× 0.30 mm). Male tergite 5 forms half-of-a-ring, bare but with a pair of long (0.175–0.20 mm) strong setae on lateral caudal edge; tergite 5 divided or at least strongly desclerotized sagittally. Sternite 5 ( Fig. 12 ) comparatively long, processes short and symmetrical with 4 pairs of thick setae. Pilosity of medial platelet rather long.

Syntergosternite 6–8 ( Fig. 8 ) long dorsally, with a large thin ventral process emerging dorsally to the ventral medial S6 part. S6 part of the complex short (as for the body axis), continued into a comparatively broad right lateral part, which continued into a ventraly and cranially directed thin process. Epandrium subdorsally with a pair of long (0.22 mm) thick setae. Male cerci large, almost as large as anal “cavity” of epandrium, cerci membranous, microsetose only. Surstylus ( Figs 9–10 ) small, of an intricate form, with a long thorn on basal lobe and with several strong setae. Postgonite ( Fig. 11 ) bifid apically with a subapical, lateral, broad based ridge (cf. Fig. 22 for Ch. ornata ) and with a rather sharp apex. Phallapodeme comparatively short with blunt apex.

Lateral caudal setae on female abdominal tergites extremely long, 0.31 mm on tergite 7. Female cerci very short, longest seta 0.017 mm.

Etymology. The specific epithet refers to the flattened male fore tarsus.

Figures

Figs 8–12. Chaetopodella latitarsis sp. n., paratype male, genitalia.8 = postabdominal sclerites, ventral view, 9 = surstylus, broadest lateral view, 10 = same, broadest ventral view, 11 = postgonite and

Figs 18–24. Chaetopodella spp. n., paratype males, genitalia. 18–20 = Ch. orientalis sp, n.: 18 = surstylus in broadest (subventral) view, 19 = postgonite and phallapodeme laterally, 20 = 5th sternite, ventral view; 21–24 = Ch. ornata sp. n.: 21 = 5th sternite, ventral view, 22= postgonite and phallapodeme laterally, 23 = surstylus, lateral view, 24 = same, ventral view. Scales: 0.05 mm for Fig. 18, 0.1 mm for Figs 19–24