Erpobdella adani, Tessler & Siddall & Oceguera-Figueroa, 2018

Tessler, Michael, Siddall, Mark E. & Oceguera-Figueroa, Alejandro, 2018, Leeches from Chiapas, Mexico, with a new species of Erpobdella (Hirudinida: Erpobdellidae), American Museum Novitates 2018 (3895), pp. 1-1 : 1-

Erpobdella adani , n. sp.

Figures 2–6

MATERIAL EXAMINED: Fifteen specimens collected in Río Fogótico, in Arcotete, Chiapas, Mexico, on January 23, 2016, by Michael Tessler, Samantha Contreras, Jairo Arroyave, and Alejandro Oceguera-Figueroa. Specimens were found beneath submerged rocks in moderately flowing water. Non-blood-feeding (macrophagous) .

DESCRIPTION: External morphology based on 15 specimens. Body strongly dorsoventrally flattened, vermiform, slender. Dorsal surface dark gray, ventral surface lighter; some specimens with three longitudinal lines of black pigment, others lacking lines. Dorsal surface smooth, without conspicuous papillae or other sensory structures; ventral surface of some specimens covered with minute papillae (figs. 2, 3). Average body length 51 mm (35–64 mm), average body width 4 mm (2–5 mm). Complete somite five annulate. Clitellum conspicuous, thick both dorsally and ventrally, comprising 15 annuli, from X b5 to XIII a2. Male gonopore at XII b2/a2, female gonopore XII/XIII; three annuli between gonopores (fig. 4). Male gonopore large, conspicuous, and surrounded by an integumental disc; female gonopore small. Three pairs of eyespots in the classic “erpobdellid” arrangement with the anterior labial pair on III and two buccal pairs at IV. Mouth wide, occupying entire anterior sucker; anterior lip blunt (fig. 5). Anus large, on XXVI/XXVII. Posterior sucker weakly developed,

Figures

FIGURES 2–6. Erpobdella adani. 2. Holotype, dorsal view. 3. Holotype, ventral view. 4. Paratype, male gonopore (black arrow), female gonopore (white arrow). 5. Paratype, ventral anterior end, anterior sucker. 6. Illustration of the male (dark gray) and female (light gray) reproductive systems, atrial horns (ah), ejaculatory duct (ed), ventral nerve cord (nc), ovary (ov); testisacs not shown.