Alloleptis Nagatomi & Saigusa

Kerr, Peter H., 2010, 2592, Zootaxa 2592, pp. 1-133 : 103

Genus Alloleptis Nagatomi & Saigusa

Figure 64

Alloleptis Nagatomi & Saigusa 1982 a: 40. Type species Alloleptis tersus Nagatomi & Saigusa 1982 a, by monotypy.

Diagnosis. Alloleptis is the only rhagionid genus with a tibial spur formula of 0:1:1. It is a small fly (body and wing length 3.8 mm), with antenna similar to those found in Bolbomyia (first flagellomere enlarged, bearing a short, two-segmented stylus). The antennal flagellomeres were lost prior to measurement and no illustration exists, however. Alloleptis may be distinguished immediately from Bolbomyia by having conspicuously setulose eyes and M 3 present. Other distinctive features include bare laterotergite, wing veins CuA 2 and A 1 join well before the wing margin (CuA 2 +A 1 long), and the wings are darkly infuscate.

Description. No specimens available for examination; character state scoring based on Nagatomi (1982 a, 1984). Head. Clypeus slightly bulbous. Scape slightly smaller than pedicel. First flagellomere enlarged bearing two-segmented stylus. Eyes conspicuously setulose; male holoptic. Palpus one-segmented or twosegmented (the original description reads "probably two-segmented (if so, basal segment short)." The illustrations of the head ( Nagatomi 1982a: 56), however, show palpus one-segmented). Mandibles absent.

Thorax. Mesonotum and scutellum dark, with long, erect setae. Wing membrane infuscate, without markings, without pterostigma. Lower calypter reduced. Upper calypter well developed, with broad curvature, lobe-like, width twice length or less. Costa apparently extends past wing tip or at least to R 5. R 2+3 nearly straight; longer than R 5, but less than twice as long. Base of R 4 –R 5 fork proximal or directly above distal end of cell dm. R 4 at base relaxed, not strongly curved, nearly straight apically. R 5 clearly longer than R 4+5 (r-m to R 4 origin); ending at wing tip. M 3 present. Cell m 3 parallel-sided at margin. Origin of CuA 1 at crossvein separating discal and basal medial cells. Length of CuA 2 vs. posterior vein of cell bm less than 1/2 length of posterior vein of cell bm. Alula with broad curvature, rounded evenly. Anal lobe well developed. Cell cu p closed (CuA 2 + A 1 vein as long or longer than CuA 2). Laterotergite bare. Subscutellum inconspicuous. Tibial spur formula 0:1:1.

Abdomen is “comparatively long and narrow” ( Nagatomi 1982a: 41). Male genitalia with epandrium simple, not containing hypandrium ventrally. Epandrium wider than long, modestly curved anteriorly. Tergite 10 absent. Hypoproct tomentose, without setae. Cercus base held underneath epandrium. Cerci directly adjacent to one another, separation distance one quarter width of cercus or less. Hypandrium fused entirely to gonocoxites. Gonocoxite smooth dorsally, without sinuous ridge leading to gonocoxal apodeme. Gonocoxal apodemes very short. Sperm sac narrow. Lateral ejaculatory processes present, integrated into sperm sac membrane. Ejaculatory apodeme moderately long, reaching slightly beyond margin, anteriorly; rod-shaped. Aedeagal tines absent. Endoaedeagal process present. Female are not known.

Larva. Unknown.

Biology. Unknown.

Literature. Illustrations of wing and head in Nagatomi (1982a); male genitalia illustrated in Nagatomi (1984).