var. scopolii

Uzunhisarcikli, M. Erkan, Güner, Ebru Doğan & Ekici, Murat, 2018, Synopsis of the genus Scrophularia (Scrophulariaceae) in Turkey, Phytotaxa 333 (2), pp. 151-187 : 160-162

15a. var. scopolii

= S. glandulosa Waldst. & Kit., Pl. Rar. Hung. 3: 238, t. 214 (1807) = S. glandulifera E.D. Clarke, Travels 1(2): 21 (1813) = S. grandidentata Ten., Fl. Nap. Suppl. 2: 69 (1819) ≡ S. scopolii var. grandidentata (Ten.) Boiss., loc. cit. (1879) = S. melissaefolia Urv., Enum. Pl. Or. 75 (1822) = S. decumbens Fisch., Mey. & Avé-Lall in Ind. Sem. Horti Petrop. 10: 58 (‘1842’ [recte, 1845]) = S. tmolea Boiss., Diagn. Pl. Orient. ser. 1(4): 66 (1844) ≡ S. scopolii var. tmolea (Boiss.) Boiss. , loc. cit. (1879) = S. grandifolia C. Koch in Linnaea 22: 707 (1849) = S. oligantha Boiss. & Heldr. in Boiss., Diagn. Pl. Orient. ser. 1(12): 33 (1833) ≡ S. scopolii var. oligantha (Boiss.) Boiss. , loc. cit. (1879) = S. puberula Boiss. & Hausskn. ex Boiss., Fl. Orient. 4: 396 (1879) = S. guestii Eig in Palestine J. Bot., Jerusalem ser. 3: 80 (1944).

Type: — Described from Caranthia, Austria (n.v.).

Specimens seen: — TURKEY. Adana: Karsantı, Torasan Mountain, Erkeç Yatağı location, 2200 m, 16 June 1977, E. Yurdakulol 10610 ( ANK) ; Afyon: Dazkırı, Maymun Mountain , streamsides, 1250 m, 22 August 1984, Z. Aytaç 1668 ( GAZI) ; Şuhut, Kumalar Mountain, Başören Village , Kıran hill, volcanic slopes, 1400 m, 13 June 1998, E. Akçiçek 2033 ( GAZI) ; Bayat, Meliktepe , ca. 1400 m, 29 June 1975, M. Vural 172 ( ANK) ; Ankara: Hacıkadın river , 28 May 1954, R. Çetik 1074 ( ANK) ; Mamak, Kıbrıs Village gorge, Dipsiz Lake-Kale , streamsides, 1100 m, 19 May 2005, S. Aslan 1935 ( GAZI) ; Kızılcahamam, Soğuksu Milli Parkı , 1200‒1400 m, 10 June 2013, M. E. Uzunhisarcıklı 2417 ( GAZI) ; Antalya: Düden , R. Çetik 3794 ( ANK) ; Amasya: Akdağ, Çalardı Plateau , 1500 m, 22 May 1977 ( ISTE 36956 ) ; Taşova, Boraboy , 1000 m, 21 May 1978 ( ISTE 39834 ) ; Artvin: Çoruh , ca. 1000 m, 19 June 1957, Davis & Hedge 29752 ( ANK) ; Şavşat, Karagöl , 1450 m, 19 September 2004, H. Altınözlü 4323 ( HUB), 1700 m, 02 July 2004, H. Altınözlü 4010 ( HUB) ; Balıkesir: Zeytinli-Kazdağ, Çeyizdere , 1200 m, 17 October 1970 ( ISTE 18766 ) ; Bitlis: Rocky , 1500 m, 23 June 1983, T. October 7486 ( GAZI) ; Bolu: Abant , 1350 m, 04 June 1977 ( ISTE 37400 ) ; Abant , 1350 m, 22 June 1979 ( ISTE 42052 ) ; SW of Abant Lake , 1500 m, 26 July 1985, E. Leblebici, Ö. Seçmen, G. Gürk ( EGE 38893 ) ; Kale, Tekneci river , streamsides, 1000 m, 08 June 1990, İ. Kılınç 1075 ( GAZI) ; Bursa: E of İznik Lake , 85 m, 15 July 1984, E. Leblebici & H. Ellikçi ( EGE 38894 ) ; NW of İznik Lake , 85 m, 25 July 1985, E. Leblebici 6156 et al. ( EGE 38891 ) ; Çorum: near the Çatak , N of Çorum, ca. 1000 m, 31 May 1965 ( ISTE 43722 ) ; Düzce: Akçakoca road, Fagus forest, 800 m, 05 June 1980, M. Aydoğdu 542 ( ANK) ; Aydınpınar, Yılankaya location, 550 m, 05 August 1982, M. Aydoğdu 781 ( ANK) ; Akçakoca, Alaplı , 1‒50 m, 29 May 2001, Ş. Yıldırımlı et al. ( HUB 27296 ) ; Akçakoca, around Paşalar Village , 40‒75 m, 11 May 2003, A. Doğrukoca 2313 ( HUB) ; Akçakoca, around Kurukavak Village , 500‒550 m, 07 June 2001, A. Doğrukoca 1273 ( HUB) ; Elazığ: Hedi (Aydınlar) Village , streamsides, ca. 850 m, 20 June 1980, H. Evren 320 ( ANK) ; Süzek Village , ca. 1250 m, 25 July 1980 Y. Altan 842 ( HUB) ; Yaylabaşı Village, Kazankaya Mountain , 1300‒2000 m, 06 August 1980, Ş. Yıldırımlı 3761 ( HUB) ; Isparta: Senirkent-Garip Village , Cedrus forest, rocky, 15 June 1984, L. Bekat 1326 ( EGE) ; Şarkıkaraağaç, Yenişarbademli , 1600‒1700 m, 30 June 1974, A. Güner 1392 et al. ( HUB) ; İstanbul: Beykoz, Beykoz Konakları , 100 m, 10 April 1999, H. Şağban 3044 ( HUB) ; Aydos , 22 April 1951 ( ISTE 3850 ) ; Belgrad forest , 29 April 1956 ( ISTE 4454 ) ; Belgrad forests, shady places, 19 June 1967 ( ISTE 11524 ) ; Belgrad forests, edge of Kılıçpınar Lake , 19 June 1967 ( ISTE 12267 ) ; Kurtkemeri, Ayvad Bendi road, 12 May 1971 ( ISTE 19697 ) ; Şile , seaside, 11 August 1972 ( ISTE 23088 ) ; Topuz Bendi-Gümüşdere Village , streamsides, 11 May 1970 ( ISTE 17967 ) ; Tuzla , 25 April 1897, J. Nemetz ( WU) ; Kahramanmaraş: Göksun, Kınıkköz Village, Mezdelipınar , 1600 m, 18 July 1977, B. Yıldız 1051 ( HUB) ; Binboğa Mountain, Karlı Dere Plateau , ca. 2600 m, 05 January 1952, P. H. Davis 20098 ( ANK) ; Göksun, Kocaağız Village , 1800 m, 13 June 1978, B. Yıldız 2134 ( HUB) ; Göksun, Çardak, Fındık Village , Bostandere , 1800 m. 14 May 1978, B. Yıldız 1756 ( HUB) ; Süleymanlı, Berit Mountain, Çimen Plateau 2500 m, 11 June 1978, B. Yıldız 2043 ( HUB) ; Engizek Mountain, Ağıllıpınar location, 2300 m, 23 June 1988, H. Duman 4078 ( GAZI) ; H. Duman 4216 ( GAZI) ; Berit Mountain, Yedikardaşlar road, streamsides, 2400 m, 08 July 2015, M. E. Uzunhisarcıklı 2697 ( GAZI) ; Karabük: Keltepe , ca. 1900 m, 13 July 1984, M. Demirörs 1928 ( ANK) ; Karaman: Ermenek, Tekeçatı, Kamış river , shady places, 1400 m, 26 August 1992, M. Vural 6160 et al. ( GAZI) ; Ermenek, Kazancı Kasabası, Yeşilköy , 700 m, 12 October 1983, H. Sümbül 2533 ( HUB) ; Kars: Posof, Kol Village- Posof , ca. 1500 m, 04 September 1985, Demirkuş 3312 ( HUB) ; Kastamonu: Ilgaz, garden of Ilgaz Mountain Resort Otel , B. Bilgili 3923 ( GAZI) ; Şenova , Quercus pubescens scrub, 18 June 1977, Y. Akman ( ANK) ; Daday, Ballıdağ , ca. 1350 m, 11 June 1979, O. Ketenoğlu 697 ( ANK) ; Daday , Abies forest, ca. 1100 m, 19 June 1978, O. Ketenoğlu 698 ( ANK) ; Ilgaz Mountain , ca. 1775 m, 23 June 1981, Y. Akman 11901 et al. ( ANK) ; Ilgaz Mountain , ca. 1700 m, 17 June 1977, Y. Akman 6836 ( ANK) ; İnebolu, Özlüce Cevizli Village , ca. 100 m, 31 May 1978, O. Ketenoğlu 695 ( ANK) ; Azdavay , near water, 1500 m, 20 May 1972 ( ISTE 21785 ) ; Ilgaz pass, 1800 m, 17 July 1976 ( ISTE 35192 ) ; Küre, Uzunöz Village , 1000 m, 22 May 2013, B. Bilgili 3833 ( GAZI) ; Küre, İsfendiyar Mountains, Karacakaya , 1320 m, 12 June 1977 ( ISTE 38549 ) ; around Yaralıgöz Mountain , Fagus forest, 13 June 1990, E. Yurdakulol 3374 ( ANK) ; Tosya, Ahlat Mountain , Pinus nigra forest, ca. 1300 m, 15 June 1977, M. Keleş 6877 ( ANK) ; Tosya, Yağcılar Village, Karanlık Dere location, ca. 1500 m, 12 June 1975, M. Kılınç 3485 ( ANK) ; from Tosya , Pinus nigra forest, ca. 1400 m, 17 June 1977, Y. Akman 6839 et al. ( ANK) ; Sarıcasu , Abies forest, ca. 1000 m, 16 June 1977, Y. Akman 6838 et al. ( ANK) ; Kayseri: İncesu , 28 July 1932, W. Kotte 489 ( ANK) ; Yahyalı, Sultan Sazlığı , 07 July 1982, H. Sümbül 1438 ( HUB) ; Sultan Sazlığı , 1000 m, 23 August 1984, Y. Gemici ( EGE 38892 ) ; Kırıkkale: Delice, Büyük Avşar Village, Büyük Hemit location, Quercus scrub, 1190 m, 10 July 1990, C. Birden 1406 ( GAZI) ; Koçubaba , 100 m, 02 July 1990, A. A. Dönmez 2589 ( HUB) ; Kırklareli: Dereköy , A. Baytop ( ISTE 14130 ) ; Mahyadağ , 12 June 1968 ( ISTE 13261 ) ; Kocaeli: Gebze, Eskihisar , 09 July 1961 ( ISTE 6749 ) ; Konya: Sultan Mountains, Çakıllar üstleri, Sarıpınar location, 1800 m, 15 July 1980 ( ISTE 45458 ) ; Kütahya: Gediz, Murat Mountain , streamsides, 1850 m, 22 July 1978 ( ISTF 32436 ) ; Murat Mountain, Kesiksöğüt Plateau , 1640 m, 03 July 1976 ( ISTE 35024 ) ; Mersin: Fındıkpınarı, Çapur Gediği çevresi, 1200‒1400 m, 14 July 1987, Ş. Yıldırımlı 9923 ( HUB) ; Niğde: Hasan Mountain, Taşpınar Plateau , ca. 1900 m, P. H. Davis & R. Çetik 19016 ( ANK) ; Rize: Ardeşen-Çamlıhemşin , 100‒250 m, 26 February 1981, A. Güner 3186 ( HUB) ; Çamlıhemşin, Aşağı Kaçkar Plateau , 1400‒1800 m, 15 August 1981, A. Güner 4185 ( HUB) ; Çamlıhemşin, around Aşağı Vice , 250‒800 m, 24 February 1981, A. Güner 3125 et al. ( HUB, ANK) ; Fındıklı, Arslandere Village, Gürcüdüzü location, 550‒700 m, 27 June 1980, A. Güner 2618 ( HUB, ANK) ; Seasides , 10 m, 25 May 1985, A. Güner 6437 ( HUB) ; Çayeli, Kaptanpaşa, Çatalpaşa Village , Çürükbel location, 1370‒1450 m, 05 July 1979, A. Güner 1996 ( HUB) ; Sakarya: Hendek , in garden, 19 May 1994 ( ISTF 36360 ) ; Samsun: Akdağ, Ladik Plateau , 1300 m, 10 October 1978 ( ISTE 41632 ) ; Samsun to Ankara, 30. km., banks of garden, 627 m, 24 July 2012, M. E. Uzunhisarcıklı 2298 ( GAZI) ; Sivas: Şarkışla, Diğnendim hill, 1600‒1900 m, 27 June 2009, B. Özüdoğru 2356 ( HUB) ; Gemerek, Karababa Mountain, Karasivri hill, 2200‒2230 m, 09 June 2007, B. Özüdoğru 1220 ( HUB) ; Yıldız Mountain , Quercus scrub, rocky, 2000‒2100 m, 19 July 2014, M. E. Uzunhisarcıklı 2617 ( GAZI) ; NW slopes of Yıldız Mountain , Quercus scrub, 2000 m, 10 July 2015, M. E. Uzunhisarcıklı 2702 ( GAZI) ; Divriği-Zara, 3‒4. km., Quercus scrub, 1700 m, 09 July 2015, M. E. Uzunhisarcıklı 2700 ( GAZI). Tokat: Artova, Akraş Village , Çal hill, streamsides, ca. 1300 m, 13 June 1981, R. İlarslan 1335 ( ANK) ; Reşadiye, Çambalı Village , 25 August 1974 ( ISTE 30908 ) ; Tunceli: Ovacık-Hozat , ca. 2050 m, 22 July 1957, Davis & Hedge 31578 ( ANK) ; Hozat, Ovacık , ca. 2000 m, 14 July 1957, Davis & Hedge 31086 ( ANK) ; Munzur Mountain, Ovacık , ca. 2400 m, 18 July 1957, P. H. Davis 31375 ( ANK) ; Hozat-Ovacık , 2000 m, 14 July 1957 ( ISTE 43721 ) ; Uşak: Gürlek, Çukuroluk , meadows, 1650 m, 18 June 1978, ( ISTF 31865 ) ; Yozgat: Çekerek, Dondurmanın Başı hill, under and in clearings Pinus sylvestris forest, ca. 1400‒1500 m, 18 June 1980, R. İlarslan 1049 ( ANK) .

Flowering time:— February–September.

Habitat:— Streamsides, seaside, under and in clearings forest, Quercus forest, Cedrus forest, Abies forest, shady places, Fagus forest, roadsides, garden, meadows, rocky places, volcanic slopes. 1‒2600 m elevation.

Distribution:— Multiregional element (Irano-Turanian, Euro-Siberian and Mediterranean element).