Pachyrhinus cinereus (Casey 1888)

Benzel, Joseph & Bright, Donald E., 2021, A revision of the genus Pachyrhinus Schӧnherr 1823 (Coleoptera: Curculionidae Entiminae) in the Nearctic Region, Zootaxa 4999 (6), pp. 501-533 : 523-531

Pachyrhinus cinereus (Casey 1888)

Scythropus cinereus Casey 1888: 276 ; Fall 1901: 309.

Pachyrhinus cinereus: O’Brien & Wibmer 1982:43 .

Scythropus crassicornis Casey 1888: 276 . Fall 1901: 309. NEW SYNONYMY.

Pachyrhinus crassicornis: O’Brien & Wibmer 1982:43 .

Scythropus albidus Fall 1901: 309 . NEW SYNONYMY.

Pachyrhinus albidus: O’Brien & Wibmer 1982:43 .

Diagnosis. Pachyrhinus cinereus ( Figs. 90–95 ) is similar to P. californicus but can be distinguished by the blunt teardrop-shaped scales on the elytra ( Fig. 8 ) along with thickened and blunt recumbent or slightly appressed elongate scales interspersed along the elytral interstria ( Figs. 96–97 ). In P. californicus the scales are elongate with tapered ends and are occasionally setiform. Specimens of P. cinereus have a mottled scale color pattern as opposed the uniformly colored interstria of P. elegans . The aedeagus of P. cinereus ( Figs. 98–100 ) is identical to that of P. californicus in being highly arcuate with a blunt tip. Pachyrhinus elegans in contrast has a moderately curved aedeagus with a long, tapered tip. The spermatheca ( Fig. 101 ) is comma shaped rather than C-shaped as in P. elegans .

Description (male). Length: 5.0–7.0 mm average 6.0 mm ( Fig. 90 ); color black, red on tibia tarsi and antennae ( Fig. 91 ); scales about 1.5× long as wide with blunt tips, teardrop shaped ( Fig. 8 ), color gold, ash grey scales on underside ( Fig. 92 ), sides, and mottled spots on elytra. Funicular antennomeres III to VII conical, decreasing in size to V, increasing in size thereafter. Striae uniformly covered in scales ( Fig. 96 ); scales formed into mottled color pattern with patches of teardrop shaped gold scales 2.0× long as wide mottled with patches of ash grey scales of the same dimensions ( Fig. 97 ); grey scales most common on stria I and VII–X. Aedeagus mesally curved ( Fig. 98 ); lateral margins curved, ending in lengthened point covered by long fine setae ( Figs. 99–100 ); ostium elongate, ovate, highly arcuate, laterally emarginate, terminating in abrupt narrow point with slight ventral curvature.

Description (female). Length 5.0–9.0 mm, average 6.5 mm. Spermatheca comma-shaped ( Fig. 101 ); ramus elongate, subconical; cornu elongate, recurved, apically narrowed.

Variation ( Figs 102–105 ). Scale color is variable, ranging from ash grey ( Fig. 103 ) to gold ( Figs. 104–105 ) and orange ( Fig. 105 ). The color of the primary scales may closely match that of the secondary scales on the underside and in mottled patches on the elytra, giving the appearance of a uniform scale color ( Fig. 102 ). The more elongate scales that appear on some specimens on the interstriae show some variation in size ranging from nearly identical to the surrounding scales to elongate and slightly appressed. The scales themselves are more loosely attached to the integument than in other species of Pachyrhinus resulting in some older P. cinereus lacking most of their scales and having a “rubbed” appearance. In New Mexico, some P. cinereus are smaller ranging from 4.5 to 5.0 mm in body length, but otherwise identical.

Distribution ( Fig. 106 ). Pachyrhinus cinereus is common in mountain coniferous forests along the Rocky Mountains from New Mexico to Montana and into Alberta and British Columbia and west to the Sierra Nevada in California.

Material Examined (571 specimens). HOLOTYPE, USA, California, Tahoe Co. T. L. Casey (female) ( USNM P. crassicornis holotype, USA, New Mexico, county not listed, ( USNM). P. albidus holotype, USA, California, Lake Co. Lake Tahoe, VII-17-21-1897. (MCZ)

CANADA, British Columbia, Baynes L. V-6-1976 / B. F. & J. L. Carr ( CNCI: 2 male) ; Cawston. IV-17-1929 / W. R. S. Metcalfe ( CNCI: 2 male, 5 female) ; Cranbrook. IV-12-1922 / C. B. D. Garrett ( CNCI: 2 males, 1 female) ; Kelowna. IV-5-1927 / E. R. Buckell ( CNCI: 1 female) ; Kamloops. V-11-1937 / J. K. Jakob ( CNCI: 1 unknown) ; Elko. V-7-1958 / H. & A. Howden ( CNCI: 2 females) ; Fort Steel. V-22-1977 / B. F. & J. L. Carr (5 males) ; North

Bend. VI-6/ Hubbard & Schwartz ( USNM: 1 female) ; Oliver. V-25-1959 / L. A. Kelton ( CNCI: 8 males, 11 females, 1 unknown) ; Orofino Mtn. VI-6-1958 / H. & A. Howden ( CNCI: 1 female) ; Osoyos, V-30-1958 / B. F. & J. L. Carr ( CNCI: 1 female) ; Peachland, IV-10-1918 / W. R. S. Metcalfe ( CNCI: 2 males) ; Skookumchuck. V-30-1984 / B. F. & J. L. Carr ( CNCI: 2 males) ; Summerland. V-6-1959 / R. E. Leech ( CNCI: 1 male) ; Vernon. IV-22/ Wickham ( USNM: 1 female) ; V-7-1924 / E. R. Buckwell ( CNCI: 1 female) ; IV-24-1924 / E. P. Venables ( CNCI: 1 unknown) ;

USA, Arizona, Apache Co., Jackson Spring near Alpine. VI-4-1982 / R. W. Baumann ( CWOB: 1 female) ; Nutrioso Cr. Hwy. 180 below Alpine Divide. VI-20-1987 / R. W. Baumann, B. C. Kondratieff, Sargent, & Wells ( CWOB: 1 male) . Cochise Co., Chiricahua Mtns. , VI-3-1935 / J. N. Knull ( CNCI: 1 female, 1 unknown) ; Onion Saddle V-4-1978 / R. Gordon ( USNM: 2 females) . Coconino Co., Williams. V-20/ Barber & Schwartz ( USNM: 1 male) ; V-26 ( USNM: 1 male, 11 females) ; V-27 ( USNM: 3 males, 10 females) ; V-28 ( USNM: 1 male, 3 females) ; V-29 ( USNM: 2 males) ; V-30 ( USNM: 1 male, 1 female) ; VI-6 ( USNM: 3 females) ; VI-8 ( USNM: 1 female) ; VI-12 ( USNM: 1 female) ; San Fran Mtns. VI-8-1915 ( USNM: 1 female) ; Tusayan NF., V-7-1915 ( USNM: 2 males, 1 female) ; San Fran. Mtns. V-8-1915 / ponderosa pine, M. R. Wagner ( CUIC: 2 males, 1 female) . Flagstaff. IV-24-1916 / ponderosa pine, C. F. Korstain ( USNM: 10 males, 21 females, 1 undetermined) . V-21-1934 / F. T. Scott ( CNCI: 1 male, 3 females) . V-5-1947 / R. S. Beal ( CWOB: 1 male) . IV-27-1983 / ponderosa pine, M. R. Wagner (2 males, 1 female). VI-10-1995 / W. B. Warner ( CWOB: 2 females) ; III-7/ Barber & Schwartz ( USNM: 1 female) . Navajo Co., Winslow. V-10-1968 / ponderosa pine, P. E. Buffam ( USNM: 1 male, 2 females) ; Apache NF. V-5-1934 / ponderosa pine, U. S. Hopkins ( USNM: 7 males, 3 females) ; White Mtns. , Diamond Cr. VI-1925 / D. Duncan ( USNM: 2 females) ; Pima Co., Catalina Mtns., Hitchcock Hwy. mi 22. V-22-1958 / C. W. O’Brien & Wibmer ( CWOB: 1 male) . Yavapai Co., Lynx Cr. 10 mi, N of Prescott. IV-29-1936 / ponderosa pine, H. Morrison ( USNM: 2 females) Oak Cr. VI-14-1948 / M. W. Stone ( UCRC: 1 female) ; Prescott. III-26-2011 / long leaf pine, C. W. O’Brien & Wibmer ( CWOB: 3 males, 1 female) ; VI-19/ Barber & Schwartz ( USNM: 1 male, 1 female) .

California, Alameda Co., Berkeley. II-2-1934 / ( CNCI: 1 male) ; Arroyo Mocho ; 20 mi. S Livermore. III-2- 1953 / D. Burdick. ( EMEC: 1 female) ; V-24-1957 / ( CWOB: 1 male) ; Arroyo Mocho 10 mi S of Livermore. III-2- 1958 / P. D. Ashlock ( CWOB: 2 females) ; Arroyo Mocho , 17 mi. SE of Livermore. II-18-1967 / knobcone pine, W. Gagne ( CWOB: 1 male, 2 females. EMEC: 1 female) ; Cedar Mtn. III-3-1968 / Digger pine, J. Powell. ( EMEC: 2 males, 2 females) ; Mt. Santini. VII-3-1971 / Timberlake ( UCRC: 1 female) ; Alpine Co., Bear Valley . VI-6-1914 / R. S. Woglum ( UCRC: 4 females) ; Butte Co., Bangor. III-28- 1955 / K. S. Hagen. ( EMEC: 2 males, 1 female) ; Calaveras Co., Rail Road Flat. VI-17-1974 / G. S. Linsley. ( EMEC: 1 female) ; Chico Co., Bidwell Park. IV-14-1974 / S. R. Palten. ( EMEC: 1 female) ; Contra Costa Co., 4 mi. N of Orinda Russel Property. III-14-1966 / C. Slobodenkoff ( CWOB: 2 males, 1 female) ; Tilden Pk. Inspiration point. IV-8/ IV-18-1992 / Monterey pine, W. Middlekauff ( EMEC: 4 males, 8 females) ; El Dorado Co., Meyers . VII-3-1930 / A. T. McClay ( CNCI: 1 female) ; Placerville. III-6-1913 / E. J. B. ( USNM: 1 male) ; nr. Whitehall. V-21-1931 / E. C. Zimmerman ( USNM: 1 female) ; Lake Tahoe. VII-11-1931 / E. C. Zimmerman. ( USNM: 1 female) ; Blodgett Forest 13 mi. E of Georgetown. V-29-1971 / ( EMEC: 4 females) ; V-29-1977 / ponderosa pine. J. Yoakley. ( EMEC: 1 female) ; S Fk. American R., Hwy 50 Riverton. III- 23-1985 / R. W. Baumann ( CWOB: 5 males, 2 females) ; Blodgett Forest 14 mi. E of Georgetown. IV-27-1985 /. M. C. Whitmore. ( EMEC: 1 female) ; 2.3 mi NE Virner. III-20-1996 / L. G. Bezark ( CWOB: 5 males, 3 females) . Ogilvy Cyn. Cr. , SE of Pacific House. IV-21-1996 / L. G. & E. A. Bezark ( CWOB: 2 females) . Pacific House. III-1-1997 / L. G. Bezark ( CWOB: 2 males) ; Mormon Emigrant Trail, 2-3 mi from Sly Park. III-2-1998 / L. G. & E. A. Bezark ( CWOB: 3males, 3 females) ; El Dorado NF, N38.4623° W120.2328°. V-18-2015 / ponderosa pine, J. S. Benzel ( CSUC: 2 males, 4 females) ; 1 mi S of Meyers. V-26/ D. G. Kissing. ( USNM: 2 females) . Fresno Co., Long Meadow Cr. At Hume L. III-16-1985 / R. W. Baumann ( CWOB: 1 male) ; Humboldt Co., Forest Home. III-10-1966 / M. W. Stone ( CWOB: 1 female) ; Inyo Co., Onion Valley . VI-17-1937 / J. H. Mitchell ( CWOB: 1 male) . Coyote Ridge. VI-26-1976 / E. Giesbert ( EMEC: 1 female) . Kern Co., Mt. Pinos. VII-2-1955 / G. A. Gorelick ( CWOB: 1 female) . W of Wofford Hts. IV-28-1964 / W. Turner. ( EMEC: 2 females) . Tehachapi Mt. Co. campground. V-6-1965 / Doyer. ( EMEC: 3 females) . Lake Co. Whispering Pines. IV-18-1964 / W. Turner. ( EMEC: 1 female) ; V-19-1964 / R. F. Luck ( CWOB: 1 female) ; Lassen Co., Laufman St. 3 mi. SE Milford. V-17-1982 / J. A. Powell. ( EMEC: 4 females) ; Gold run Cr. Gold run Rd. SW Susanville. V-22-2007 / B. C. Kondratieff, R. W. Baumann. ( CSUC: 1 female) ; Marin Co., Bodega Bay . II-6-1988 / J. K. Robertson ( EMEC: 4 males, 9 female) ; Mariposa Co., Yosemite. VI- 7-1928 / A. T. McClay ( CNCI: 1 male) ; 5-28-1931 / D. W. Clancy ( UCRC: 1 female) ; Big Meadow. III-20-1969 / D. L. Tiemann. ( EMEC: 1 female) ; Modoc Co., NW of Canby. VI-24-1960 / E. L. Schlinger ( UCRC: 2 females) S Fk. Parker Cr. FRd. 31. V-22-2007 / D. Leatherman. ( CSUC: 2 males, 2 females) ; Mono Co., Blanco’s Corral, White Mtn. VI-27- 1953 / H. Nalahibarra ( UCRC: 1 female) ; Monterey Co., Carmel. III-13-1915 / J. F. Brimley ( CNCI: 3 male) . Prunedale. I-1965 / Monterey Pine, C. Kohler ( CWOB: 1 male) ; Nevada Co. , IV-13-1963 / L. V. & T. L. Erwin ( UCRC: 1 male) ; Alta Sierra. VI-14-1961 / Ponderosa pine, H. Howden ( CNCI: 1 male) ; Grass Valley . V-13-1971 ( UCRC: 3 males) ; 5 mi. SW grass Valley. V-6-1980 / ponderosa pine and Jeffery pine, J. T. Doyen and J. K. Liebherr ( EMEC: 1 male, 7 females) ; Donner Picnic Area . N39°22’41 W120°09’42. V-19-2015 / ponderosa pine, J. S. Benzel ( CSUC: 1 female) ; Orange Co., Meadow Valley , VII-11-1924 / A. C. Davis ( CNCI: 1 female) . Placer Co., Forest Hill. IV-1898 ( USNM: 2 males, 1 female) ; Dutch Flats. II-28-1909 ( UCRC: 3 females) ; Flume, Dog Bar Rd. V-27-1989 / R. Gordon ( USNM: 2 males, 1 female) ; Plumas Co., VI-30/ E. J. Blum. ( CNCI: 1 female) ; 7 mi N of Chester on Juniper L. Rd. VI-4-1964 / T. L. Erwin ( CWOB: 1 female) ; Benner Cr. VI-5-1964 / T. L. Erwin ( CWOB: 1 female) ; NW Chester on Benner Cr. VI-1-1965 / T. L. Erwin ( CWOB: 2 males) ; VI-11-1965, ( USNM: 2 females) ; Squirrel Cr. 8 mi E of Quincy. V-15-1982 / G. W. Urlich. ( EMEC: 1 male, 1 female) ; Humbug Cr. 3 mi. W of Portola. V-16-1982 / J. A. Chemsak. ( EMEC: 8 males, 8 females) ; San Bernardino Co., Near Fallsvale, San Gorgonio Mtn. VI-23-1964 / J. D. Pinto ( UCRC: 1 female) ; Running Springs. VI-9-1965 / J. T. Doyen ( EMEC: 1 female) . USA, CA, Bear Lake . VII- 5-1932 / M. W. Stone ( UCRC: 1 female) ; Seven Oaks. V-13-1962 / H. Ewing ( UCRC: 1 female) ; San Luis Obispo Co., Paso Robbles. Shoemaker ( USNM: 3 males, 4 females) ; Santa Barbara Co., San Marcos Pass. IV-14-1960 / E. I. Schinger and J. C. Hall ( UCRC: 1 female) ; Orcutt. III-5/ III-25-1966 / A. J. Kissler ( USNM: 7 males, 6 females) ; Mt. Hamilton. V-6-1957 / D. Burdick ( CWOB: 1 male) ; Santa Clara Co., Los Gatos. III-30-1962 / T. L. Erwin ( USNM: 1 male) ; Mt. Hamilton. VII-14-1967 / knobcone pine, W. Gagne ( CWOB: 1 male) ; Shasta Co., Platina. V-27-1977 / B. F. & J. L. Carr ( CNCI: 1 female) ; Siskiyou Co. V-29-1911 / F. W. Nunenache ( USNM: 1 female) ; Sonoma Co., Rhododendron St. Pk. IV-15-1955 / D. J. Burdick ( EMEC: 2 females) ; 4 mi W of Plantation. IV-6-1957 / D. Burdick ( CWOB: 4 male) ; Mt. St. Helena. V/ L. W. Saylor ( USNM: 5 males, 6 females) ; Stanislaus Co., Adobe Cr. II-26- 1948 / R. F. Smith. ( EMEC: 1 male) ; Tuolumne Co., Strawberry , V-20-1993 / B. F. & J. L. Carr ( CNCI: 1 male) ; Trinity Co., 15 mi. SW Hay Fk. IV-17-1962 / ponderosa pine, R. W. Clabaugh ( CWOB: 1 male) ; Tulare Co., Sequoia NP. VI-1932. F. T. Scott ( USNM: 1 female) ; Tuolumne Co., Sand Flat. VI-11-1930 / D. W. Clancy ( UCRC: 1 female) ; VI-26-1931. H. E. Hinton ( USNM: 1 female) ; Twain Harte. II-5-1967 / M. R. Lundgrin ( EMEC: 1 male) ; Ventura Co., Chuchupate Rgr. Sta. base. Frazier Mtn. V-1-1955 / Jeffery Pine, C. W. O’Brien & Wibmer ( CWOB: 1 female) ; Mt. Pinos, Campground 1. VIII-19-1970 / Jeffery pine, R. A. Medeveo ( UCRC: 1 female) ; Sierra foothill field sta. 6 mi. N of Smartville. V-4-1980 / J. A. Powell. ( EMEC: 2 males) .

Colorado, Boulder Co., Boulder. VI-8-1961 / W. R. M. Mason ( CNCI: 1 female) ; Chaffee Co., Dreney Gulch. VI-24-1999 / ponderosa pine, D. Leatherman. ( CSUC: 1 female) ; Elbert Co., Jct. CR 94/CR77. VI-27-1995 / ponderosa pine, D. Leatherman. ( CSUC: 2 females) ; Las Animas Co., Trinidad lakes St. Park N side picnic area. V-18- 2012 / D. Leatherman. ( CSUC: 1 male) ; Mesa Co., Colorado National Monument. V-11-1949 / Pinyon Pine ( USNM: 3 males) ; Weld Co., E of CR 115 at WY line. IV-18-1994 / limber pine, D. Leatherman. ( CSUC: 3 males, 9 females, 1 undetermined) .

Idaho, Clearwater Co., Orofino. IV-25-1967 / ponderosa pine, G. C. Franc ( USNM: 2 females); Lewis Co. Winchester. V-1-1924 / M. C. Lane ( USNM: 3 males, 7 females) .

Montana, Lewis & Clark Co., Helena. IV-21-1907 / W. M. Mann ( USNM: 2 males) .

New Mexico, McKinley Co., 1–2 mi. S of Ft. Wingate. VI-6-1988 / pinyon pine, R. Turnbow ( CWOB: 1 male, 2 females) ; IV, ( USNM: 1 female) ; Otero Co., Cloudcroft , V-20-1967 / H. F. Howden (2 male, 1 female) ; Santa Fe Co ., Santé Fe. VI-5, ( USNM: 23 males, 20 females) ; San Juan Co. , Aztec. V-5, C. F. Baker ( USNM: 1 female) ; San Miguel Co., Las Vegas HS. III-8/ Barber & Schwartz ( USNM: 1 female) .

Nevada, Washoe Co., Jct. Hwy. 15 & 28. VI-27-1962 / C. W. O’Brien & Wibmer ( CWOB: 2 males, 4 females) ; Whites Cr., Whites Cr. Trailhead. VI-16-2009 / B. C. Kondratieff, R. W. Baumann. ( CSUC: 1 female) ; Thomas Cr. Rd. 49. VI-16-2009 / B. C. Kondratieff, R. W. Baumann. ( CSUC: 1 female) .

Oregon, Baker Co., Pine Cr. VI-8-1939 / J. H. Baker ( USNM: 1 female) ; V-1-1940 ( USNM: 1 female) ; V-5- 1940 ( USNM: 2 female) ; V-27-1945 ( USNM: 1 male, 1 female) ; Columbia Co., Clatskanie. IV-14-1936 / Douglas fir, K. Grey & J. Sohuh ( USNM: 6 males, 4 females) ; Corvallis. V-3-1936 / G. Ferguson ( USNM: 1 male) ; Cook Co., Ochoco NF. Wiley Cr. Camp. VI-20-1998 / G. R. Ballmer ( UCRC: 1 female) ; Grant Co., Silvies R. Highway 359. S Seneca. S Rd. 37. V-21-2013 / B. C. Kondratieff. ( CSUC: 2 males, 1 female) ; Hood River Co., Hood River . Gresham. V-21/ C. C. Sperry ( USNM: 1 female) ; V-21/ Hubbard & Schwartz ( USNM: 8 males, 12 females) ; Jackson Co., Butte Falls . V-17-1956 / F. P. Larson ( USNM: 1 female) ; Josephine Co., 6.5 mi S of Cave Jct. V-27-1980 / knobcone pine, R. L. Westcott ( CWOB: 2 females) ; Lake Co., Fremont NF, N42.1993° W120.6161°. V-15-2015 / white pine, J. S. Benzel ( CSUC: 7 males, 6 females) ; Klamath Co., 2 mi. NW of Chiloquin. V-17-1975 / ponderosa pine, R. L. Westcott ( CWOB: 1 female) ; Tillamook Co., Sand Lake . V-12-1940 / K. M. Fender ( USNM: 4 males, 1 female) ; Umatilla Co., Battle Mtn. V-27-1984 / B. F. & J. L. Carr ( CNCI: 2 males) ; Union Co., Kamila. VI-10-1925 / M. C. Lane ( USNM: 1 male) ; Blue Mtns. VI-4-1933 / M. C. Lane ( USNM: 4 males, 2 females) ; Cold Spring, Blue Mtns. T. McLane ( USNM: 1 male, 1 female) ; Wapinita Pass. V-5-1938 / K. Grey & J. Senuh ( USNM: 1 female) ; Perry , VI-6-1984, B. F. & J. L. Carr ( CNCI: 1 male) .

Utah, Carbon Co., 415 mi. N of Price. 143. V-20-1968 / D. G. Kissinger ( USNM: 1 male, 1 female) ; Garfield Co., 4.5 mi S Panguitch , hwy. 143. IV-23-2005 / R. Turnbow ( CWOB: 4 females) ; Dixie NF, Canaan Peak. V-27- 2003 / S. M. Clark & C. R. Nelson ( CWOB: 2 female) ; Lonesome Beaver campground, VII-30-1968, H. F. Howden ( CNCI: 1 female) . Kane Co., ponderosa Grove Rec Area. IV-20-2005, R. Turnbow ( CWOB: 1 female) ; San Juan Co., 1.5 mi. W of Utah line on US 160. V-20-1968 / D. G. Kissinger ( USNM: 2 females) ; Devil’s Cyn. nr. Campground off Hwy 191. V-29-2013 / gambel oak/ W. Crenshaw, D. Leatherman & B. C. Kondratieff. ( CSUC: 1 female) ; Sevier Co., Water Cyn. Above jct. Quitchupah Cr. IV-25-2002 / R. W. Baumann & Clark ( CWOB: 2 males) .

Washington, Clallam Co., Sol Duc Hot Springs. VI-11-1933, M. C. Lane ( USNM: 1 female) ; Columbia Co., Lewis Pk., Blue Mtns. VII-11-1933 / M. C. Lane ( USNM: 2 males, 2 females) ; Kittitas Co., Vicinity of Cle Elum. VI-25-2009 / M. Huether ( CWOB: 1 female) ; Spokane Co., Spangle. / C. V. Piper ( USNM: 1 female) ; Skagit Co., Cypress Is. V-27-1979 / M. Whitmore ( EMEC: 1 female) ; Thurston Co., Tenino. / Hubbard and Schwartz ( USNM: 1 female) .

Wyoming, Albany Co., 1.7 mi N of Easterbrook Campground. V-19-2008 / R. Turnbow ( CWOB: 3 males, 5 females); Park Co. , [Yellowstone] National Park. VII-30/ Hubbard & Schwartz ( USNM: 2 females) .

Biology. The biology of this species is similar to P. californicus . Adults and larvae feed mainly on ponderosa pine and Douglas fir.

Remarks. Smaller specimens of P. cinereus occur in New Mexico and Arizona. These were originally described as P. crassicornis ( Figs. 107-112 ) by Casey (1888) from a single specimen collected at Fort Wingate, New Mexico. Pachyrhinus crassicornis was distinguished by a shortened prothorax and a highly arcuate antennal scape only barely surpassing the posterior margin of the eye. Examination of the holotype found it to have a scape and prothorax of the normal length of P. cinereus , reaching to the anterior margin of the prothorax ( Fig. 108 ). In addition, specimens from the type locality were between 5–7mm in length. Specimens of P. cinereus from other locations occasionally showed smaller size and more arcuate antenna.

Pachyrhinus albidus ( Figs. 113-118 ) was described from a single specimen collected around Lake Tahoe, the same type locality as that of P. cinereus . The holotype bears a close resemblance to a P. cinereus with closely matching scale colors, but with only isolated scales of the secondary scale color ( Fig. 113 ). The scale morphology is identical with that of P. cinereus ( Fig. 117 ).

Figures

FIGURE 106. Pachyrhinus cinereus distribution map. Black circles represent examined specimens. Red shading delimits approximate range.

FIGURES 90–95. Habitus photos of female Pachyrhinus cinereus, holotype, female. 90. Dorsal view. 91. Lateral view. 92. Ventral view. 93. Frontal view. 94. Elytral vestiture. 95. Labels.

FIGURES 6–9. Elytral Scales. 6. Pachyrhinus elegans. 7. Pachyrhinus californicus. 8. Pachyrhinus cinereus scale. 9. Polydrusus impressifrons Gyllenhall 1834.

FIGURES 96–101. 96. Pachyrhinus cinereus left elytron. 97. P. cinereus stria. 98. P. cinereus aedeagus lateral view. 99. Ventral view. 100. Dorsal view. 101. Spermatheca.

FIGURES 102–105. Pachyrhinus cinereus vestiture variation. 102–103. Monterey Co., California. 103. Monterey Co., California. 104: Ravalli Co., Montana. 105: El Dorado Co., California.

FIGURES 107–112. Pachyrhinus crassicornis, holotype, male. 107. Dorsal view. 108. Lateral view. 109. Ventral view. 110. Frontal view. 111. Elytral vestiture. 112. Labels.

FIGURES 113–118. Pachyrhinus albidus, holotype, female. 113. Dorsal view. 114. Lateral view. 115. Ventral view. 116. Frontal view. 117. Elytral vestiture. 118. Labels.