Anagallis monelli var. genuina Pau

Nualart, Neus, Soriano, Ignasi, Prieto, David Pérez & Ibáñez, Neus, 2021, Catalogue and typification of the Moroccan taxa published by Carlos Pau, Phytotaxa 519 (1), pp. 1-94 : 10-14

Anagallis monelli var. genuina Pau in Font Quer, Iter Marocc. 1927 nº 470 (1928), in sched., nom. inval.

= Lysimachia monelli (L.) U.Manns & Anderb. subsp. monelli

Notes:—On the label there is a handwritten observation by Pau about the variability of this specimen, indicating that it is similar to plants from Cádiz ( Spain). The name of the variety should be considered invalid according to Art. 24.3 ( Turland et al. 2018).

1,3 Anagallis monelli var. speciosa Pau & Font Quer in Font Quer, Iter Marocc. 1930 nº 506 (1932), in sched.

=§ Lysimachia monelli (L.) U.Manns & Anderb. subsp. monelli

Ind. loc.:—“Hab. in schistosis, pr. Sidi Ali (Beni Aros), ad 300 m. alt.; 6 majii”

Lectotype (designated here):—In schistosis, pr. Sidi Ali (Beni Aros), 300 m., 6 May 1930, Font Quer 506-1930 (lectotype BC 98173 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98173 .jpg]; isolectotypes BCN 12667 !, GDA 35627 [photo!], MA 91527 !, MPU 006718 [photo!], S 09-44942 [photo!])

Notes:—The GDA specimen was labelled as syntype by Vizoso in 2013. The BC specimen was identified as Anagallis monelli L. by Martinell in 2004, without indicating any infraspecific taxa; the same criterion as used in APD (2020), which synoymizes this variety to Lysimachia monelli . Based on the size and shape of the leaves of the type material, it should be synonymized under subsp. monelli . Vizoso (2018) mentioned as type the specimen MA 91526!, but it corresponds to a locality (Jebel Mago) and collection date (June 1930) different from those of the original material.

1,2,3 Andryala gracilis Pau in Bol. Soc. Aragonesa Ci. Nat. 17: 130 (1918)

= Andryala integrifolia L.

Ind. loc.:—“Tifasor; y fueron recogidas desde el día 10 al 30 de Julio del año 1915”

Holotype:— Bisanor , Melilla (Tifasor?) [m. N. Ibáñez], June 1916 [sphalm. for 1915], Aterido s.n. ( BC 36093 ! [https://www.ibb.csic.es/herbari/ JPEG / BC36093 .jpg])

Notes:—The label on the specimen came from “Museu de Catalunya. Herbari” and was written by a transcriber, which would explain the error both in the locality and date. There is no “Bisanor” in Melilla: it must have been an erroneous transcription for Tifasor (as already indicated in the label by Ibáñez, curator of the Herb. BC). Moreover, Aterido was in Melilla only in 1915 for military service, and by 1916, he was working in Barcelona (see González-Bueno & Gomis 2007).

1,2,3 Andryala × humilis Pau in Font Quer, Iter Marocc. 1927 nº 728 (1928), in sched., pro sp.

= § Andryala × faurei Maire [= Andryala integrifolia L. × A. arenaria (DC.) Boiss. & Reut. ]

Ind. loc.:—“ Hab. in umbrosis Yebel Dahar, pr. Axdir (Beni Urriaguel), 300 m. alt., solo calcareo; 9 majii” Lectotype (designated by Talavera & Talavera 2015: 304):—In umbrosis Yebel Dahar, pr. Axdir (Beni Urriaguel), 300 m, 9 May 1927, Font Quer 728-1927 (lectotype BC 36140 ! [https://www.ibb.csic.es/herbari/ JPEG / BC36140 . jpg]; isolectotypes BCN 17373 [photo!], BM 001124968 [photo!], GDA 3571 [photo!]*, MA 139322 !, MA 139323 !, MPU 006365 [photo!]*)

Notes:—The BM specimen was labelled as isotype by Ferreira in 2013, and the GDA specimen as syntype by Vizoso in 2013. The GDA and MPU specimens were cited as isotypes by Talavera & Talavera (2015) and Dobignard (2009), respectively. Montserrat (2002b), Fennane (2014a), and APD (2020) synonymized the name by Pau under A. integrifolia , but according to Talavera & Talavera (2015) it should be considered a hybrid between A. integrifolia and A. arenaria , due to the original material having intermediate characters between these two species. The correct name of this hybrid is A. × faurei , which was published by Maire (1926) and thus have priority.

1 Andryala integrifolia nothovar. vidalii Pau in Vidal, Asoc. Esp. Progr. Ci. Congr. Cádiz 6: 151 (1928), pro var.

= Andryala × caballeroi Font Quer in Cavanillesia 7: 83 (1935) [= Andryala integrifolia L. × A. rothia Pers. ]

Ind. loc.:—“-”

Lectotype (designated by Talavera & Talavera 2015: 304):—Dar Riffien, June 1926, Aterido s.n. ( MA 139360 [photo!] [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00139360])

Notes:—The specimen was labelled as lectotype by Talavera in 2015. Talavera & Talavera (2015) consider this taxon an interspecific hybrid between Andryala integrifolia and A. rothia , whose correct name is A. × caballeroi ( Font Quer 1935) .

2,3 Anthemis tuberculata var. tetuanensis Pau ex Maire in Bull. Soc. Hist. Nat. Afrique N. 22(8): 296 (1931)

= Anthemis abylaea (Font Quer & Maire) Oberpr.

Ind. loc.:—“Tetuan”

Lectotype (designated by Oberprieler 1998: 229):— Maroc: Tetuan, sites bas, April 1929, Mas Guindal s.n. (lectotype MPU 001112 [photo!] [https://herbier.umontpellier.fr/hv/wakka.php?wiki=PagePrincipale/iframe& action=detail&ordre=p&codebarre=MPU001112]; isolectotypes BC 84440!*, FI 000790 [photo!], FI 000791 [photo!], G 00022020 [photo!]*, MA 127040*, MPU 001113 [photo!], PH 00003506 [photo!]) Notes:—Oberprieler (1998) considers as isolectotypes the BC, G, and MA specimens. The BC, G, and MPU specimens were identified as Anthemis abylaea and labelled as isolectotypes by Oberprieler in 1997. Sennen distributed this original material in his exsiccata Plantes d’Espagne 1929, number 6980.

1,2,3 Anthyllis hosmariensis Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 315 (1924a)

= Anthyllis vulneraria subsp. maura (Beck) Maire

Ind. loc.:—“Beni Hozmar, en los peñascos de Gorgues”

Lectotype (designated here):—Beni Hosmar, 8 May 1921, Pau s.n. (lectotype MA 65010 ! [http://colecciones.rjb. csic.es/#cardAdv.php?CatalogNumber=MA-01-00065010]; isolectotype MA 65011 !)

Notes:—The specimens were identified as A. vulneraria subsp. maura by Cullen in 1960. Pau labelled the lectotype as “ A. tingitana var. hosmariensis ” and the isolectotype as A. hosmariensis . When he published this new taxon he used the specific rank. The chosen lectotype is the one with the leaves in better condition.

2,3 Anthyllis maura var. ajmasiana Pau in Font Quer, Iter Marocc. 1928 nº 205 (1929), in sched.

=§ Anthyllis vulneraria subsp. maura (Beck) Maire

Ind. loc.:—“Hab. in saxosis calc montis Kalaa, supra Xauen, 1000 m. alt.; 13 majii”

Lectotype (designated here):—In saxosis calc montis Kalaa, supra Xauen, 1000 m, 13 May 1928, Font Quer 205- 1928 (lectotype BC 15013 !* [https://www.ibb.csic.es/herbari/ JPEG / BC15013 .jpg]; isolectotypes B *, BCN 17657 [photo!], BM *, MA 65026 !, MPU 006405 [photo!], G 00022363 [photo!], S 14-35054 [photo!])

Notes:—In Carrasco (1975) this taxon appears as Anthyllis vulneraria var. ajmasiana Pau. The BC and B specimens were cited as types by Rothmaler (1941) and the BM specimen by Cullen (1976). The MA specimen was identified as A. vulneraria subsp. maura var. maura by Cullen in 1961. In APD (2020) this taxon is accepted at the subspecies rank [ A. vulneraria subsp. ajmasiana (Pau) Raynaud & Sauvage ], but according to Benedí (2000) and Fennane (2007) subsp. maura also includes plants with yellow flowers (the diagnostic character of the variety of Pau).

1 Anthyllis polycephala var. ballii Pau in Font Quer, Iter Marocc. 1929 nº 266 (1930a), in sched.

= Anthyllis polycephala Desf.

Ind. loc.:—“Hab. in rupibus calc. montis Targsunt (Bocoia), 400 m alt.”

Lectotype (designated here):—In rupibus calc. montis Targsunt (Bocoia), 400 m, 16 May 1929, Font Quer 266- 1929 (lectotype BC 802798 ! [https://www.ibb.csic.es/herbari/ JPEG / BC802798 .jpg]; isolectotypes BCN 72438 [photo!], MA 65240 !, MPU 006934 [photo!])

Note:—The BC specimen was identified as Anthyllis polycephala by Benedí in 1993.

1,2,3 Anthyllis tangerina Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 315 (1924a)

= Anthyllis vulneraria subsp. maura (Beck) Maire

Ind. loc.:—“Colinas bajas de Tánger ”

Lectotype (designated here):— Tanger (Marabut), April 1921, Pau s.n. (lectotype MA 65035 ! [http://colecciones. rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00065035]; isolectotype MA 65036 !)

Notes:—The MA specimens were identified as A. vulneraria subsp. maura by Cullen in 1960. Pau labelled the lectotype as “ A. vulneraria β tangerina ” and the isolectotype as A. tangerina . When he published the taxon he used the specific rank. The chosen lectotype is the specimen with the largest sample.

1,2,3 Anthyllis vulneraria var. arenicola Pau in Font Quer, Iter Marocc. 1930 nº 345 (1932), in sched.

= Anthyllis vulneraria subsp. maura (Beck) Maire

Ind. loc.:—“Hab. pr. El Araix, versus Nador, in arenosis, ad 20 m. alt.; fl. 19 mart., fr. 16 apr.”

Lectotype (designated here):— Pr. El Araix, versus Nador, 20 m, fl. 19 March 1930, fruct. 16 April 1930, Font Quer 345-1930 (lectotype BC 98151 ! [https://www.ibb.csic.es/herbari/ JPEG / BC98151 .jpg]; isolectotypes BC 840080 !, BC 908894 !, BCN 74437 [photo!], GDA 33315 [photo!], MA 65001 !, MPU 006742 [photo!], MPU 006741 [photo!], S 14-35052 [photo!])

Notes:— The MPU and GDA specimens were labelled as syntypes by Mathez and Vizoso in 1982 and 2013, respectively. The MA specimen was identified as A. vulneraria subsp. maura by Cullen in 1960. The BC specimen, with more samples and labelled with the original exsiccata label, is selected as lectotype .

1,2,3 Antirrhinum glutinosum var. africanum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 571 (1928), in sched.

= Antirrhinum hispanicum Chav.

Ind. loc.:—“ Hab. in rupium arenacearum, juxta Bu-Meziat (Atlante rhiphaeo), 1600 m. alt., r. r.; 29 junii” Lectotype (designated here):—In rupium arenacearum, juxta Bu-Meziat (Atlante rhiphaeo), 1600 m, 29 June 1927, Font Quer 571-1927 (lectotype BC 44107 ! [https://www.ibb.csic.es/herbari/ JPEG / BC44107 .jpg]; isolectotypes BCN 17244 !, MA 110647 !, MPU 006554 [photo!]*)

Note:— The BCN specimen was labelled as syntype and the MPU specimen cited as isotype by Dobignard (2009) .

1 Antirrhinum murale f. leptophylla Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 368 (1924a)

= Antirrhinum tortuosum Bosc ex Vent.

Ind. loc.:—“Tetuán”

Holotype:—Tetuan, 6 May 1921, Pau s.n. ( MA 110547 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00110547])

1,2 Antirrhinum orontium f. leiocarpum Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 367 (1924a)

= Misopates calycinum (Lam.) Rothm.

Ind. loc.:—“De Tánger a Ceuta ”

Type:—not seen

Note:—The form name was published as A. orontium var. calycinum f. leiocarpum .

1,2,3 Antirrhinum tortuosum var. hosmariense Pau in Font Quer, Iter Marocc. 1930 nº 596 (1932), in sched.

= Antirrhinum tortuosum Bosc ex Vent.

Ind. loc.:—“Hab. in rupibus montis Gorgues (Beni Hosmar), ad 700 m. alt.; 30 majii”

Lectotype (designated here):—In rupibus montis Gorgues (Beni Hosmar), 700 m, 30 May 1930, Font Quer 596- 1930 (lectotype BC 808514 ! [https://www.ibb.csic.es/herbari/ JPEG / BC808514 .jpg]; isolectotypes BC 98431 !, BCN 73723 [photo!], MA 110760 !, MPU 005263 [photo!], MPU 005264 [photo!], S 10-19857 [photo!]) Notes:— Carrasco (1975) records this taxon as “ Antirrhinum majus f. hosmariense ” Pau. The selected lectotype is the BC specimen with flowering samples, because the corolla size is the diagnostic character of this variety.

1,2,3 Antirrhinum vidalianum Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 572 (1928), in sched.

= Antirrhinum tortuosum Bosc ex Vent.

Ind. loc.:—“Hab. in fissuris rupium schist., pr. Zarkat (Atlante rhiphaeo), 1200 m. alt.; 16 julii”

Lectotype (designated here):—In fissuris rupium schist., pr. Zarkat (Atlante rhiphaeo), 1200 m, 16 July 1927, Font Quer 572-1927 (lectotype BCN 17243 ! [http://www.ub.edu/cedocbiv/herbarivirt/70304]; isolectotypes BC 44163 !, BM 000930593 [photo!], G 00396557 [photo!]*, MA 110768 !)

Notes:— The BCN specimen was labelled as syntype. The G specimen was cited by Dobignard (2009) as isotype. The BCN specimen is selected as lectotype due to it being complete and in better condition than the BC one .

1 Aquilegia vulgaris var. riffienensis Pau in Broteria Ser. Ci. Nat. 5(3): 114 (1936)

= Aquilegia vulgaris subsp. cossoniana (Maire & Sennen) Dobignard & D.Jord.

Ind. loc.:—“Cultivada de semillas, proporcionadas por el Sr. Hno. Sennen, y procedentes de Tidiguín (Rif)”

Holotype:— s.l., s.d., s.c. ( MA 38824 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01- 00038824])

Notes:—The original label handwritten by Pau has some taxonomical notes concerning this variety, but there is no indication of locality, collector or date. Based on examination of the original material, it should be synonymized under A. vulgaris subsp. cossoniana .

1,2,3 Arabis alpina var. commutata Pau & Font Quer in Font Quer, Iter Marocc. 1928 nº 138 (1929), in sched.

= Arabis alpina subsp. caucasica (Willd.) Briq.

Ind. loc.:—“Hab. in saxosis calc. montis Magot, 1300 m. alt., 21 majii”

Lectotype (designated by Talavera & Velayos 1992: 147):—In saxosis calc. montis Magot, 1300 m, 21 May 1928, Font Quer 138-1928 (lectotype MA 48667 ! [http://colecciones.rjb.csic.es/#cardAdv.php?CatalogNumber=MA-01-00048667]; isolectotypes B 10 0154784 [photo!], BC 2903 !, BCN 38806 !, BM 000583665 [photo!], GDA 28915 [photo!], MPU 006423 [photo!], MPU 006424 [photo!], Z 000112196 [photo!])

Note:— The BCN and GDA specimens were labelled as syntypes, the latter one by Vizoso in 2013 .

1,2,3 Arabis caucasica var. leiopoda Pau in Font Quer, Iter Marocc. 1929 nº 197 (1930a), in sched., nom. nud.

= Arabis alpina subsp. caucasica (Willd.) Briq.

1,2,3 Arabis decumbens var. brachypoda Font Quer & Pau in Font Quer, Mem. Acad. Ci. Artes Barcelona 22(18): 346 (1931b)

= Arabis pubescens subsp. decumbens Ball

Ind. loc.:—“Hab. in cacumine montis Lexhab, ad 2.150 m alt.”

Lectotype (designated here):—In rupibus umbrosis calc., cacuminis Montis Lexhab, 2150 m, 25 June 1930 fl., 21 July 1930 fr., Font Quer 260-1930 (lectotype MA 48584 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00048584]; isolectotypes BC 802945 !, BCN 73020 !, GDA 28940 [photo!], MPU 002804 [photo!], MPU 002803 [photo!])

Notes:— The GDA specimen was labelled as syntype by Vizoso in 2013. The MA specimen, which includes flower and fruit samples, is selected as lectotype, instead of the BC specimen that only has flowering plants .

1,2,3 Arabis leptopoda Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 236 (1928), in sched.

= Arabis josiae Jahand. & Maire

Ind. loc.:—“ Hab. in saxosis, lacisque silvaticis montis Iguermalez (Atlante rhiphaeo), 1950 m. alt.; 7 junii” Lectotype (designated here):—In saxosis, lacisque silvaticis montis Iguermalez (Atlante rhiphaeo), 1950 m, 7 June 1928, Font Quer 236-1928 (lectotype MA 48513 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00048513]; isolectotypes BC 2978 !, BCN 17301 !, BM 000583666 [photo!], GDA 28946 [photo!], MPU 006331 [photo!], MPU 006332 [photo!])

Notes:— The BCN and GDA specimens were labelled as syntypes, the latter one by Vizoso in 2013. The MA specimen is selected as lectotype, as it is the only one with mature siliquae .

1,2,3 Arabis pubescens var. leucanthemifolia Pau & Font Quer in Font Quer, Iter Marocc. 1927 nº 235 (1928), in sched.

≡ Arabis pubescens subsp. leucanthemifolia (Pau & Font Quer) Maire

Ind. loc.:—“ Hab. in rupestribus arenaceis cacuminis Yebel Lerz (Atlante rhiphaeo), 2000 m. alt.; 12 junii” Lectotype (designated here):—In rupestribus arenaceis cacuminis Yebel Lerz (Atlante rhiphaeo), 2000 m, 12 June 1927, Font Quer 235-1927 (lectotype BC 2982 ! [https://www.ibb.csic.es/herbari/ JPEG / BC2982 .jpg]; isolectotypes BCN 17302 !, MA 48589 !, MPU 004660 [photo!], MPU 004659 [photo!])

Note:— The BCN and MPU specimens were labelled as syntype, those at MPU by Madaule, Durand & Loup in 2004 .

1,2 Arabis saxatilis f. latiorifolia Pau in Font Quer, Iter Marocc. 1929 nº 198 (1930a), in sched., nom. nud.

= Arabis nova subsp. iberica Rivas Mart. ex Talavera

Note:—The accepted name follows Montserrat (2002c).

2,3 Arbutus unedo var. tangerina Pau in Mem. Real Soc. Esp. Hist. Nat. 12: 358 (1924a)

= Arbutus unedo L.

Ind. loc.:—“Yebel Quebir”

Holotype:—Djebel Kebir, May 1921, Pau s.n. ( MA 89629 ! [http://colecciones.rjb.csic.es/#cardAdv. php?CatalogNumber=MA-01-00089629])