Hymenocephalus papyraceus, Jordan & Gilbert, 1904

Iwamoto, Tomio, Nakayama, Naohide, Shao, Kwang-Tsao & Table, Hsuan-Ching Ho, 2015, Synopsis of the Grenadier Fishes (Gadiformes; Teleostei) of Taiwan, Proceedings of the California Academy of Sciences 62 (3), pp. 31-126 : 76

Hymenocephalus papyraceus Jordan and Gilbert, 1904

Figure 12 .

Hymenocephalus papyraceus Jordan and Gilbert, 1904:614. fig. (holotype USNM 50935, 147 mm TL; Sagami Bay, Japan, 120–265 fm [219–485 m], Albatross sta. 3697).— Okamura, 1970:56–58, pl. XVII, text-fig. 24 (1 spec., 95 mm TL, Pacific coast s. Japan).— Sazonov, 1993:117–121 (fig.) (1 spec., East China Sea; 800–826 m).

Hymenocephalus (Papyrocephalus) papyraceus : Gilbert and Hubbs, 1920:539 (included as one of three species in new subgenus).

Hymenocephalus papiraceus : Okamura in Masuda et al., 1984:94, pl. 344–I (species name misspelled; compiled).

MATERIAL EXAMINED.— NET: NMMB-P 9121 (1, 25.7 HL), Da-xi.

DISTINGUISHING FEATURES.— 1D II,10; P i14; V 11; GR-I (outer/inner) 0+16/5+17, GR-II 5+17/4+16. Orbit 23% HL, 1.4 into postorbital; interorbital 35%; internasal width 23%; suborbital 19%; postorbital 54%; orbit to preopercle angle 54%; upper jaw 47%; barbel 3.5%; 1D-2D interspace 60%; P length 70%; V length 62%; length outer V to A origin 54%; pre-anal length 144%; length outer gill slit 27%. Head relatively deep and broad; orbit not included in dorsal profile; nape high, beginning slightly behind posterior edge of orbit. Snout with high median nasal ridge, flexible, pointed tip projecting slightly beyond mouth. Posterior end of preopercle ridge narrowly chisel-shaped (or flathead shovel-shaped). Barbel small but distinct. Head color mostly black, but dorsal head bones and snout transparent. Entire trunk to over A origin deeply and heavily peppered; remainder of body covered with large melanophores. Ground color medium brown; A base black anteriorly. Gular and BR membranes black. Attains about 150 mm TL

REMARKS.— Hymenocephalus papyraceus is apparently a rare species, having been recorded only three times previously: the holotype (and only type specimen), a small specimen in poor condition reported by Okamura (1970), and the East China Sea specimen of Sazonov (1993). The Taiwan specimen is the first record from outside Japanese waters.

Figures

FIGURE 12. Hymenocephalus papyraceus Jordan and Gilbert, 1904. NMMB-P9121, 25.7 mm HL, preserved, photo reserved.