Amarochara effeminata, Assing, 2010

Assing, V., 2010, A revision of Amarochara of the Holarctic region. IV. Three new species from China, a new synonymy, additional records, and an updated key to species (Coleoptera: Staphylinidae: Aleocharinae: Oxypodini), Linzer biologische Beiträge 42 (2), pp. 1139-1154 : 1146-1154

Amarochara effeminata nov.sp. ( Figs 17-25 )

T y p e m a t e r i a l: Holotype [probably slightly teneral]: " China: Yunnan, Dali Bai Aut. Pref., Wuliang Shan , 20 km NW Weishan, 25°19'58''N, 100°07'59''E, 1900 m, creek valley, litter & old flood debris sifted, 17.IX.2009, leg. M. Schülke [CH09-58] / Holotypus Amarochara effeminata sp.n. det. V. Assing 2010" (cAss).

D e s c r i p t i o n: Body length 3.0 mm. Habitus as in Fig. 17 . Coloration: body almost uniformly reddish-brown; legs reddish; antennae dark-brown, with antennomeres I- II reddish.

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Head ( Fig. 18 ) posteriorly without distinct neck, weakly transverse and of subquadrate shape, lateral margins subparallel in dorsal view; punctation very fine; interstices with shallow, but distinct microsculpture ( Fig. 19 ); eyes moderately large and moderately convex, shorter than the distance from posterior margins to posterior carina of head; genal carina fine, but fully visible in lateral view. Maxillary palpus moderately slender, preapical palpomere little more than twice as long as wide. Antennae massive; antennomere I less than twice as long as wide and with pronounced dorso-apical furrow; II shorter and more slender than I, about twice as long as broad; III shorter than II, distinctly coniform, and approximately 1.5 times as long as wide; IV disc-shaped, approximately 3 times as wide as long; V-X slightly wider than IV, of subequal width, of gradually increasing length, and decreasingly transverse, X less than twice as wide as long; XI of ovoid shape and almost as long as the combined length of VIII-X ( Fig. 20 ).

Pronotum ( Fig. 18 ) 1.17 times as wide as long and 1.36 times as wide as head, posterior angles weakly marked; lateral margins convex, maximal width slightly behind the middle; punctation relatively dense and fine; interstices glossy, with shallow microsculpture visible only at higher magnification ( Fig. 21 ); pubescence relatively short and depressed.

Elytra at suture 0.83 times as long as pronotum; posterior margin distinctly sinuate near posterior angles; punctation relatively dense, moderately fine (much coarser than that of pronotum), and defined ( Fig. 22 ); interstices with extremely shallow, almost obsolete microsculpture visible only at high magnification, particularly near posterior margin ( Fig. 22 ); pubescence depressed. Hind wings fully developed. Mesoventrite with pronounced median carina; process of mesoventrite long and apically narrowly truncate, reaching more than halfway between mesocoxae. Metatarsomere I elongated, approximately as long as the combined length of metatarsomeres II-IV.

Abdominal tergites III-V with moderately deep, densely and coarsely punctate, but not carinate anterior impressions; tergites VI-VII anteriorly with conspicuous coarse and striate punctation; remainder of tergal surfaces with very sparse and fine punctation; tergites III-VI without, tergites VII-VIII with very shallow microsculpture visible only at high magnification; interstices very glossy on all tergites.

: tergite VIII posteriorly weakly convex; posterior margin of sternite VIII broadly convex and with dense long marginal setae; median lobe of aedeagus slender, 0.36 mm long; ventral process much longer than basal capsule, straight in lateral view ( Figs 23- 24 ); paramere 0.33 mm long, not distinctly modified, apical lobe small and with four long setae ( Fig. 25 ).

: unknown.

E t y m o l o g y: The specific epithet (Latin, adjective) refers to the small aedeagus.

C o m p a r a t i v e n o t e s: Based on the similar external characters, well as the similar male sexual characters Amarochara effeminata is undoubtedly closely related to A. schuelkei . It is readily distinguished from all other congeners known from China particularly by the reddish-brown coloration of the body alone. From A. schuelkei , it is additionally separated by the less broad pronotum, the more pronouced microsculpture and the denser punctation of the forebody, the somewhat smaller eyes, the longer genal carinae, the pronounced median carina of the mesoventrite, and particularly by the distinctly smaller median lobe and parameres of the aedeagus.

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D i s t r i b u t i o n a n d b i o n o m i c s: Thetypelocality is situated in Yunnan, China, to the south of Xiaguan. The holotype was sifted from leaf litter and old flood debris in a stream valley at an altitude of 1900 m.

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4. Key to the Amarochara species of the Holarctic region

In the first comprehensive part of the revision of Amarochara , a key to the species of the genus known from the Holarctic region at that time was provided. Since then, several species have been newly described or syonymized. In order to account for the recent changes, a new key is given below. One doubtful species, Amarochara tingitana JARRIGE 1952 , which was originally described from Morocco, is not included because its external and sexual characters are unknown. The holotype is lost and no additional material has become available ( ASSING 2002a).

References to illustrations in previous papers are abbreviated as follows: A02a = ASSING (2002a); A02b = ASSING (2002b); A03 = ASSING (2003); A07 = ASSING (2007).

1 Head posteriorly without a distinct neck.............................................................................2

- Head posteriorly with a distinct neck. Eastern Palaearctic region.....................................25

2 Relatively large species, 3.0-5.0 mm. Abdominal tergite VI with distinct anterior impression (though this impression is somewhat shallower than those of tergites III-V). Elytral hind margins near posterior angles strongly sinuate. Mesoventrite with median carina. Aedeagus and spermatheca as in A02a: Figs 71-72, 74. W-Palaearctic (A02a: Map 6)...................................................................................... A. forticornis (LACORDAIRE)

- Abdominal tergite VI at most with very indistinct anterior impression. Elytral hind margins near posterior angles not or only weakly sinuate. Mesoventrite usually without, with very short, only in one species from Spain with long carina. Genitalia different...............................................................................................................................3

3 W-Palaearctic species..........................................................................................................4

- E-Palaearctic or Nearctic species......................................................................................14

4 Smaller species, 2.2-3.2 mm. Genal carina completely absent. Eyes very large and bulging, distinctly projecting from lateral outline of head. Head of ± distinctly subquadrate shape ...............................................................................................................5

- Larger species. Genal carina present, though often very fine. Eyes smaller and/or less bulging, usually distinctly shorter than postgenae in dorsal view and weakly projecting from lateral contours of head; the latter without or with weakly marked posterior angles........6

5 On average larger, 2.2-3.2 mm. Antennae relatively slender, at least approximately 0.7 mm long; antennomeres IX and X less than twice as wide as long (A02a: Fig. 5 ). Aedeagus and spermatheca as in A02a: Figs 8-10 . Palaearctic region from Western Europe eastwards to Siberia, North Korea, and China (A02a: Maps 1-2)............................. ........................................................................................................ A. umbrosa (ERICHSON)

- Body length 2.2 mm. Antennae approximately 0.6 mm long; antennomeres IX and X approximately twice as wide as long. Aedeagus unknown. Spermatheca similar to that of A. umbrosa . Species of doubtful status, possibly synonymous with A. umbrosa . Yemen................................................................................................... A. rougemonti PACE

6 Large species, length approximately 4.5 mm. Coloration distinctive: elytra bicoloured, with the anterior and lateral margins blackish and with the posterior and interior two thirds of each elytron rufous, thus forming a large reddish spot; head, pronotum, and abdomen blackish. Whole body without microsculpture. Elytra with dense and coarse punctation.: median lobe of aedeagus very long and slender and, at base of ventral process, with distinct median process (A02a: Figs 122-123). Morocco................................ ........................................................................................................... A. freyi (BERNHAUER)

- Coloration different, elytra not bicoloured. Head and pronotum either of paler coloration (except for dark specimens of A. cribripennis ) or head with distinct microsculpture.: median lobe of aedeagus without median process at base of ventral process and/or less slender..................................................................................................7

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7. Larger species, 3.8-5.3 mm. Pronotum large in relation to head, 1.15-1.25 times as wide as long and approximately 1.4 times as wide as head or even wider. Antennomere III as long as II. Punctation of anterior impressions of abdominal tergites III-V very coarse and dense. Eastern Mediterranean............................................................................8

- Smaller species. Pronotum less than 1.4 times as wide as head, only in one species from NW-Africa of similar relative width. Antennomere III often shorter than II. Punctation of anterior impressions of abdominal terga III-V less coarse and less dense. Absent from the Eastern Mediterranean, except for one species with fine punctation in anterior impressions of tergites III-V................................................................................10

8 Body larger, 4.5-5.3 mm. Head and pronotum blackish brown.: aedeagus of distinctive morphology, ventral process of median lobe not narrowed at base (ventral view), with small process at base of ventral process (A03: Figs. 38-39). Southwestern Anatolia................................................................................................ A. wunderlei ASSING

- Body smaller, 3.8-4.5 mm. Head and pronotum usually of paler coloration. Male sexual characters different. Central southern Anatolia and Middle East.............................9

9 Median lobe of aedeagus in lateral view with distinct median process at base of ventral process (A02a: Figs 58-59). Central southern Anatolia to Lebanon (A02a: Map 5).............. ............................................................................................................. A. siculifera ASSING

- Median lobe of aedeagus with longer ventral process and in lateral view without median process at base of ventral process (A02a: Figs 62-63). Israel (A02a: Map 5) ........... ................................................................................................................ A. inermis ASSING

10 Coloration of body uniformly blackish. Head and pronotum with distinct microsculpture. Punctation in impressions of tergites III-V and near anterior margin of tergite VI coarse and dense. Spermatheca as in A02a: Fig. 68. Tunisia.......... A. caeca ASSING

- Colour of body at least partly paler. Head and pronotum with distinct microsculpture only in a pale-coloured species.........................................................................................11

11 Antennae more massive and longer (at least approximately 1.2 mm) (A02a: Fig. 69). Metatarsomere I at least slightly longer than the combined length of the three following tarsomeres (A02a: Fig. 70). Punctation in the anterior impressions of tergites III-V coarse and dense. Western Mediterranean...............................................................12

- Antennae less massive and shorter than 1.2 mm. Metatarsomere I shorter than the combined length of the three following tarsomeres..........................................................13

12 Mesoventrite with median carina. Antennae with much more transverse antennomeres; antennomere V approximately 3 times as wide as long; eyes much shorter than postgenae in dorsal view; pronotum smaller and less broad, approximately 1.3 times as wide as head; elytra shorter, at suture approximately 0.75 times the length of pronotum. Legs much shorter; approximate length of metatibia and metatarsus 0.62 mm and 0.51 mm. Spermatheca as in A02a: Fig. 78. Northern Spain..................................... A. carinata ASSING

- Mesoventrite without median carina. Antennae with less transverse antennomeres (A02a: Fig. 69); antennomere V approximately twice as wide as long; eyes almost as long as postgenae in dorsal view; pronotum large and broad, approximately 1.4 times as wide as head; elytra shorter, at suture approximately 0.85 times the length of pronotum. Legs much longer; approximate length of metatibia and metatarsus 0.74 mm and 0.63 mm, respectively (A02a: Fig. 70). Sexual characters unknown. Morocco.............. ........................................................................................... A. crassicornis (QUEDENFELDT)

13 Head and pronotum with shallow, but distinct microsculpture. Colour of body yellowish-brown to pale-brown. Antennae shorter and less massive (A02a: Fig. 50). Abdomen (especially the anterior tergites) with sparser and finer punctation. Aedeagus as in A02a: Figs 44-45. Widespread from Anatolia in the southeast to France and Great Britain in the west and northwest (A02a: Map 3) ............................. A. bonnairei (FAUVEL)

- Head and pronotum without distinct microsculpture. Body colour usually darker. Antennae longer and more massive (Fig. 57). Abdomen (especially the anterior tergites) with denser and coarser punctation. Aedeagus as in A02a: Figs 52-53. Corsica and Sardinia ...................................................................... A. cribripennis MULSANT & REY

14 E-Palaearctic species.........................................................................................................15

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- Nearctic species.................................................................................................................22

15 Small species with fine genal carinae. Abdomen with anterior impressions of tergites III-V shallow and posteriorly ill-defined; sternites III-V with weak anterior constriction; punctation distinctive: on tergites III-VI weakly granulose; on tergites III- V rather dense and well-defined, but not very coarse, not coarser and denser in anterior impressions than on remainder of tergal surface; on tergite VI slightly coarser and denser than on tergites III-V, and on tergite VII finer and sparser than on tergites III- VI. Pubescence of forebody short, dense, and decumbent. Pronotum almost as wide as elytra. Antenna slender (similar to that of A. umbrosa ). Spermatheca as in A02a: Fig. 28. Northern India............................................................................ A. sororcula CAMERON

- Anterior impressions of abdominal tergites III-V deeper and with coarser punctation than posterior tergal portions. Pubescence of forebody longer, less dense, and usually ± suberect. Antenna - except for the widespread A. umbrosa - more massive....................16

16 Small species with completely absent genal carinae and slender antennae (A02a: Fig. 5 ). Head of subquadrate shape and with ± distinctly marked posterior angles. Aedeagus and spermatheca as in A02a: Figs 8-10 . Widespread Palaearctic species; in the Eastern Palaearctic eastwards to West Siberia, North Korea, and China (A02a: Maps 1-2)............... ....................................................................................................... A. umbrosa (ERICHSON)

- Mostly larger species. Genal carina present, sometimes reduced, but rudiments usually visible at least near occiput. Antennae more massive. Head often with weakly marked or indistinct posterior angles. Genitalia different..............................................................17

17 Large and pale-coloured species. Genal carina pronounced. Pronotum very large, approximately 1.4 times as wide as head. Punctation in anterior impressions of tergites III-V and near anterior margin of tergite VI very dense and coarse; punctation on remainder of tergal surfaces finer than in anterior impressions of tergites III-V, but conspicuously coarse, somewhat granulose, and dense. Spermatheca of characteristic morphology (A02a: Fig. 112). Japan............................................. A. densepunctata ASSING

- Colour of body in most species darker. Punctation of posterior areas of abdominal tergites III-V less coarse, less dense, and not granulose. Absent from Japan....................18

18 Smaller and more slender species, <3.5 mm; pronotum relatively slender, 1.13-1.25 times as wide as head.: median lobe of aedeagus smaller, approximately 0.32 mm long, and shaped as in A02a: Figs 21-22 ; spermatheca as in A02a: Fig. 24 . Northern India, Nepal................................................................................. A. heterogaster CAMERON

- On average larger species, 3.0- 4.1 mm; pronotum broader in relation to head, except for one species from Taiwan.: median lobe of aedeagus larger, at least approximately 0.36 mm long, and of different shape. Species from China and Taiwan................................19

19 Elytra much (ca. 1.4 x) broader than pronotum. Pronotum slender, approximately 1.05 times as wide as long and 1.2 times as wide as head, and with extremely dense fine punctation; interstices narrower than diameter of punctures. Forebody with distinct microsculpture and with suberect pubescence. Metatarsus approximately 0.7 times as long as metatibia; metatarsomere I shorter than combined length of II-IV. Whole abdomen with dense punctation. Aedeagus and spermatheca as in A02a: Figs 104-105, 107. Taiwan (A02a: Map 8).. ............................................................. A. formosana ASSING

- Elytra only slightly (1.20-1.25 x) broader than pronotum. Pronotum broader, distinctly transverse, at least 1.1 times as wide as long and at least 1.3 times as wide as head. Punctation of pronotum not extremely dense, interstices at least as wide as diameter of punctures ( Fig. 11 ). Elytra dark-yellowish. Forebody without distinct microsculpture and with depressed pubescence. Metatarsus at least approximately 0.8 times as long as metatibia; metatarsomere I at least as long as combined length of II-IV. Abdomen with less dense punctation. Genitalia different. China.............................................................20

20 Body of almost uniformly reddish-brown coloration and slighly smaller, approximately 3 mm long.: aedeagus small, 0.36 mm long, and shaped as in Figs 23-24 ; paramere small, 0.33 mm long ( Fig. 25 ). China: Yunnan ................................... A. effeminata nov.sp.

- Body blackish-brown to blackish, with the elytra sometimes paler,> 3 mm long.: aedeagus either distinctly longer (0.45 mm) or of completely different shape.................21

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21 Eyes smaller and less bulging, approximately as long as the distance between posterior margin of eye and posterior carina in dorsal view ( Fig. 2 ). Antennomeres IV and V disc-shaped, approximately 3 times as broad as long ( Fig. 3 ). Maxillary palpus moderately slender, preapical palpomere approximately 2.5 times as long as broad. Posterior margin of elytra near postero-lateral angles strongly sinuate ( Fig. 2 ). Forebody with less fine and less dense punctation.: median lobe of aedeagus longer, approximately 0.45 mm long, and with conspicuously long and slender apical portion ( Figs 5-6 ). China: Yunnan ..................................................................... A. schuelkei nov.sp.

- Eyes larger and strongly bulging, longer than distance between posterior margin of eye and posterior carina in dorsal view ( Fig. 9 ). Antennomeres IV and V not disc-shaped, barely twice as wide as long ( Fig. 10 ). Maxillary palpus long and slender, preapical palpomere approximately 4 times as long as broad. Posterior margin of elytra near postero-lateral angles weakly sinuate ( Fig. 9 ). Forebody with finer and denser punctation ( Fig. 11 ).: median lobe of aedeagus shorter, approximately 0.38 mm long, and with shorter and less slender apical portion ( Figs 12-13 ). China: Zhejiang ........... ............................................................................................................. A. hamulata nov.sp.

22 Head and pronotum with conspicuously coarse microsculpture, almost completely matt (A07: Figs 2 , 20 ).: median lobe of aedeagus as in Figs 8-12 . Eastern Canada: New Brunswick ............................................................................................ A. formicina ASSING

- Head and pronotum with or without very shallow microsculpture, shiny. Male sexual characters different............................................................................................................23

23 Colour of body ± uniformly brown or ferrugineous, only abdominal tergite VI and sometimes head and antennae ± infuscate. Antennae shorter, less than 0.75 mm long. Eyes shorter and less bulging, postocular region at least 1.5 times as long as eyes in dorsal view. Punctation of abdomen very fine, only in anterior impressions of tergites III-V somewhat more distinct.: median lobe of aedeagus and paramere as in A07: Figs 15-18 .: spermatheca as in A02a: Fig. 38. U.S.A.: Iowa; Canada: New Brunswick ............................................................................................ A. inquilina (CASEY)

- At least head and abdomen distinctly darker. Antennae longer than 0.75 mm. Eyes larger and more bulging, distinctly projecting from lateral contours of head; postocular region less than 1.5 times as long as eyes in dorsal view. Punctation of abdomen very distinct...............................................................................................................................24

24 Larger species, 3.0- 3.8 mm. Coloration on the whole usually darker, pronotum at most only slightly paler than the dark-brown to blackish head; antennae not distinctly bicoloured. Antennae longer and with less transverse antennomeres; antennomeres VII-IX approximately twice as wide as long (A02a: Fig. 37). Pronotum relatively large, 1.25-1.30 times as wide as head. Elytra relatively short, at suture approximately 0.75 times as long as pronotum. Legs distinctly longer, length of metatarsus even in small specimens at least 0.37 mm long; first metatarsomere very elongate, as long as the combined length of the three following tarsomeres or nearly so (A02a: Fig. 35). Abdomen with more clearly defined and less dense punctation, and with pronounced shine.: aedeagus of completely different shape, with much longer and more slender ventral process (A02a: Figs 29-30).: spermatheca smaller (A02a: Fig. 32). Indiana, Kansas, Georgia ................................................................................ A. fenyesi BLATCHLEY

- Smaller species, 2.8-3.2 mm. Pronotum pale-brown to brown, distinctly paler than the blackish head; antennae distinctly bicoloured, dark brown to blackish, with yellowish to ferrugineous basal antennomeres. Antennae shorter and with more transverse antennomeres; antennomeres VII-IX almost 3 times as wide as long. Pronotum relatively smaller. Elytra longer, at suture> 0.80 times as long as pronotum. Legs much shorter, length of metatarsus approximately 0.30 mm; first metatarsomere of similar length as in A. umbrosa , shorter than the three following tarsomeres. Abdomen with less defined and much denser punctation, and therefore with reduced shine.: median lobe of aedeagus with much shorter and broader ventral process (A02a: Figs 39-40).: spermatheca larger (Fig. 42). Kansas ..................................... A. brevios ASSING

25 Abdominal tergites III and IV with one, tergite V with two conspicuous transverse series of dense and coarse punctures in posterior half. Small species. Nepal (A02a: Map 7)............................................................................................ A. seriepunctata ASSING

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- Abdominal tergites III-V without distinct transverse series of dense and coarse punctures in posterior half. Larger species........................................................................26

26 Median lobe of aedeagus and parameres strongly modified. Ventral process of median lobe short and small in relation to basal portion, in lateral view with short process or obtuse projection near base of ventral process; internal sac with several strongly sclerotized structures. Apical lobe of paramere distinctly flattened (A02a: Figs 84, 85, 95).....................................................................................................................................27

- Aedeagus and paramere not strongly modified. Ventral process of median lobe longer in relation to basal portion, in lateral view without short process or obtuse projection near base of ventral process; internal sac with weakly sclerotized structures. Apical lobe of paramere not distinctly flattened...........................................................................29

27: apex of ventral process of aedeagus in lateral view obliquely truncate and in ventral view keeled, internal structures distinctive (A02b: Figs 1-2 ); apical lobe of paramere flattened, transverse, and apically truncate (A02b: Fig. 3 ). Nepal................ A. sequax ASSING

-: ventral process of aedeagus slender in lateral view and not keeled; internal structures of aedeagus and apical lobe of paramere of different shape.............................28

28 Ventral process of median lobe longer, in lateral view near base of ventral lobe with short and acute process; sclerotized structures of aedeagus of distinctive shape (A02a: Figs 82-83, 86); apical lobe of paramere not transverse, apically rounded (A02a: Figs 84-85); spermatheca as in A02a: Fig. 87. Nepal (A02a: Map 7).................... A. loebli PACE

- Ventral process of median lobe shorter, in lateral view near base of ventral lobe with obtuse projection; sclerotized structures of aedeagus different (Figs 92-93); apical lobe of paramere transverse, apically truncate (A02a: Figs 94-95). China (Sichuan, Shaanxi) (A02a: Map 8).......................................................................................... A. armata ASSING

29 Antennae much longer, antennomeres VII-X approximately 1.5 times as wide as long. Eyes larger, approximately as long as postocular region in dorsal view. Posterior halves of abdominal tergites III-V with distinct, though not very dense punctation. Aedeagus and spermatheca as in A02a: Figs 100-101, 103). Nepal and Taiwan (A02a: Map 8)................................................................................................. A. megalops ASSING

- Antennae distinctly shorter, antennomeres VII-X more than twice as wide as long. Eyes smaller, somewhat shorter than postocular region in dorsal view. Posterior halves of abdominal tergites III-V with very fine and sparse punctation. Aedeagus and spermatheca as in A02a: Figs 96-97, 99. China: Sichuan, Shaanxi, Yunnan ......................... ................................................................................................................. A. wrasei ASSING

Acknowledgements

I am indebted to the colleagues listed in the material section for the loan of material under their care. In particular, I am grateful to Michael Schülke for the generous gift of the holotypes of A. schuelkei and A. effeminata , as well as to Jon Cooter, Hereford, for providing additional material. Benedikt Feldmann (Münster) proof-read an earlier version of the manuscript.

Zusammenfassung

Ischnopoda turfanensis PACE 1998 nov.syn. wird mit Amarochara umbrosa (ERICHSON 1837) synonymisiert. Amarochara schuelkei nov.sp. ( China: Yunnan), A. hamulata nov.sp. ( China: Zhejiang) und A. effeminata nov.sp. ( China: Yunnan) werden beschrieben und abgebildet. Für mehrere Arten, davon eine unbeschrieben, werden weitere Nachweise gemeldet. Eine aktualisierte Bestimmungstabelle der Arten der Holarktis wird erstellt. In der Holarktis ist die Gattung derzeit mit 30 beschriebenen Arten vertreten.

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References

ASSING V. (2002 a): A taxonomic and phylogenetic revision of Amarochara THOMSON. I. The species of the Holarctic region (Coleoptera: Staphylinidae, Aleocharinae, Oxypodini). — Beiträge zur Entomologie, Keltern 52 (1): 111-204.

ASSING V. (2002 b): A new species of Amarochara THOMSON from the Himalayas (Coleoptera: Staphylinidae, Aleocharinae, Oxypodini). — Linzer biologische Beiträge 34 (1): 297-298.

ASSING V. (2003): New species and records of Staphylinidae from Turkey (Insecta: Coleoptera: Staphylinidae). — Entomologische Blätter 98 (2002): 153-177.

ASSING V. (2007): A revision of Amarochara of the Holarctic region. III. A new species, a new synonym, and additional records (Coleoptera, Staphylinidae: Aleocharinae, Oxypodini). — Zootaxa 1411: 25-32.

PACE R. (1991): Aleocharinae dello Yemen (Coleoptera, Staphylinidae). — Bollettino del Museo civico di Storia naturale di Verona 15 (1988): 125-150.

PACE R. (1998): Aleocharinae della Cina: parte II (Coleoptera, Staphylinidae). — Revue suisse de Zoologie 105 (2): 395-463.

PAĝNIK G. (2006) A revision of the world species of the genus Tachyusa ERICHSON, 1837 (Coleoptera, Staphylinidae: Aleocharinae). — Zootaxa 1146: 1-152

SMETANA A. (2004): Staphylinidae, subfamily Aleocharinae, pp. 353–494. — In: LÖBL I. & A. SMETANA (eds): Catalogue of Palaearctic Coleoptera. II. Hydrophiloidea – Histeroidea – Staphylinoidea. — Stenstrup, Apollo Books: 942 pp.

Author´s address: Dr. Volker ASSING

Gabelsbergerstr. 2

D-30163 Hannover, Germany

E-mail: [e-mail]

Figures

Figs 17-25: Amarochara effeminata nov.sp. (holotype): (17) habitus; (18) forebody; (19) median dorsal portion of head; (20) antenna; (21) median dorsal portion of pronotum; (22) posterior portion of left elytron; (23) median lobe of aedeagus in lateral view; (24) ventral process of median lobe of aedeagus in ventral view; (25) paramere. Scale bars: 17: 1.0 mm; 18, 20: 0.5 mm; 19, 21-25: 0.1 mm.

Figs 1-8: Amarochara schuelkei nov.sp. (holotype): (1) habitus; (2) forebody; (3) antenna; (4) abdomen; (5) median lobe of aedeagus in lateral view; (6) ventral process of median lobe of aedeagus in ventral view; (7) paramere; (8) apical portion of paramere. Scale bars: 1: 1.0 mm; 2-4: 0.5 mm; 5-8: 0.1 mm.

Figs 9-16: Amarochara hamulata nov.sp.: (9) forebody; (10) antenna; (11) postero-median portion of pronotum and antero-median portion of elytra; (12-13) median lobe of aedeagus in lateral and in ventral view; (14) apical portion of median lobe of aedeagus in lateral view; (15) paramere; (16) spermatheca. Scale bars: 9: 0.5 mm; 10-16: 0.1 mm.